Individual site joint state/signal probability matrix.

pc probability that an empty site is occupied (colonized) next year.
pn probability that a site is re-occupied next year if fledging is not successful.
ps probability that a site is re-occupied next year if fledging is successful.
pu(h) probability that fledging is successful if access to the site is unrestricted.
pr(h) probability that fledging is successful if access to the site is restricted.
pu and pr depend on the size of the arctic hare population h, which is assumed known at the time restriction decisions are made.