TABLE 3 in Contributions to the knowledge of the genus Trichodrilus Claparède (Lumbriculidae, Clitellata) with the description of new groundwater species from Spain

TABLE 3. Selected characters and measurements of Trichodrilus stygodytes sp. nov., T. tenuis Hrabě and T. moravicus Hrabě, comparing original descriptions with measurements made by authors on the NMP type material and on populations inhabiting groundwaters of Spain. Abbreviations: L, length; W, width; Ø, diameter; ind, individual; ant, anterior; post, posterior; sg, segment. Roman numbers indicate the order of the segment. Note (1): measured by the authors on mated individuals or/and unmated with sperm in sperm funnels. CharactersT. stygodytes n.sp. (Spanish type collection)T. stygodytes n.sp. (Býčí Skála) (1)T. moravicus (type collection— Říčka River) (1)T. tenuis (original description)T. tenuis (4 individuals, type collection) (1)T. tenuis (Spanish collection)Body Ø (mm) (s. X)0.17–0.300.30–0.40 (0.56 in Hrabe 1938)0.21–0.31 (0.26–0.33 in Hrabe 1938)0.190.17–0.26 (wholemounts)0.20–0.35 (whole-mounts and dissected)Dorsal anterior chaetae (L, µm)c.50–84 dorsals=0.80– 1.0 ventrals77–112 dorsals = 0.80–0.96 ventrals60–96 dorsals = 0.8–1 ventrals64–7740–6841–103 (dorsals= 0.7– 0.9 ventrals)Ventral anterior chaetae (L, µm)(55) 62–10090–13068–10664–7753–7161–107 (grow from II–VIII)ClitellumX–XII (anterior XIII)X–anterior XIIIX–anterior XIIIIndistinct–X–XII (XIII)Pharyngeal glandsV–VIIIIV or V–VIIIV–VIIIIV–VIIV–VIIV (V)–VII or VIIIDouble annulationFrom III or IVFrom IIIFrom IIIFrom IIIFrom IIIFrom III or IV (V)Atrium, total L x ampulla Ø (µm)80–121 x 50–80 (petiolate, ampulla spherical-oval)79–127 x 43– 80) (petiolate, ampulla oval)110–164 x 53–91 (petiolate, ampulla spherical-oval)73 x 5889–98 x 36–56 (not petiolate, ampulla oval)63–112 x 32–75 (not petiolate, ampulla oval)Atrial duct L x Ø (µm)Short, cylindrical 30–40 x 17–30Short, cylindrical 32–54 x 16–27 (10–17 at pore)Short, cylindrical, 31–51 x 20–27 (to 12–17 at pore)–c. 30–40 x 24c. 23–42 x 16–26PenisShort, salient or indistinctShort, salient or indistinctShort, conical (salient in 2 individuals)No penial chamberShort, extruded through the pore in one specimen.Short, may be extruded.Male pore on a porophore (L, µm)Yes, roundconical (15–20 high)YesYes, roundconical (32 µ high)Yes, minute, conicalYes, conical c. 25 µm highYes, c. 25 µm highAtrial musculature (µm)2–8, to 10 at the vasa junction2–5, to 7 at the vasa junctionc. 2–3c. 22–3c.2–3Total atrium length to body Ø ratio0.3–0.50.3–0.4c. 0.5–0.3–0.50.2–0.4Prostatic cells (height, µm)To 47–80, in clustersTo 43–77, in usually welldefined clustersTo 80–94Long, apicalTo 47–61To (34) 54–104 diffuse or in few large clustersVasa deferentia junctionAt the base of the ampullaAt the base of the ampullaSubapical or median–SubapicalMedian to subapicalVasa def. Ø (µm)8–13 Penetrate in XI10–24 Penetrate in XI14–24 (to 8, close to atrium). Penetrate in XIPenetrate in XI9–16 Penetrate in XI6–14 Penetrate in XISpermathecal ampullae (L x W, µm)c.50–120 x 50–70 In same sg. as the pore or passing into next sg.115–144 x 70–108 In same sg. as the pore or passing into next sg.100–247 x 60–130 In same sg. as the pore or passing into next sgAmpullae extend into the next sg175–232 x 58– 74 In the same sg, or passing into next sg.88–255 x 30–120 In the same sg. or passing into next sg.Spermathecal ducts (L x W, µm)40–70 x 10–15 (21–30 Ø at ectal vestibule)70–74 x 10–20 (20–33 Ø at ectal vestibule)105–160 x 15–30 (25–35 Ø at ectal vestibule)Short, distinctly separated duct34–48 x 9–11 (21 Ø ectally)28–72 x 9–12 (21–27 Ø ectally)SitesCaves and resurgence, Sta. Eufemia-Ereñozar karst (Bizkaia, Spain)Cave Býčí Skála specimens, Moravian Karst (Czech Republic, Hr 1026, 1028).All measurements in Říčka River, Moravian Karst (Czech Republic)Moravian Karst and Bohemia in Czech RepublicType collection, National Museum of Prague (Hr 1517-9 #4. Hr 1517-7 #1)Cave rivers, resurgences and wells (Spain) ......continued on the next page