Acoustic stability in hyrax snorts: vocal tightrope-walkers or wrathful verbal assailants?

The source-filter theory proposes that information on caller properties is communicated through acoustic qualities, as physical state and performance ability are reflected in the voice. Vocal stability, manifested through harshness is especially intriguing, and has rarely been explored although harsh sounds are prevalent in nature. Male rock hyraxes (Procavia capensis) produce loud complex calls that we term songs. Only the calls of older, socially dominant males, include a harsh sound termed snort. As snorts are the rarest element in songs, we hypothesized that high quality snorts are difficult to produce, and that their quality consists in the ability to maintain smoothness throughout this low-pitched, harsh call. We quantified harshness by measuring periodicity deviations and expected to find a link between social parameters (residence, rank, and weight) and the ability to produce longer, smoother snorts. In addition, we presumed that if calls are used as vocal contests, conspecifics...