Comparison and test for deviation from neutrality of the RCPE and ZE sequence variability among species.

Π: Average number of nucleotide differences per site between two sequences (equations 10.5 or 10.6 in [90]), its standard deviation is the square root of its sampling variance (equation 10.7 in [90]). Θ-W: Watterson estimator of variability (equation 1.4a in [91]) per site (equation 10.3 in [90]). Tajima's D: Suggested in [92] for testing whether all variation is selectively neutral. Fu and Li's D: Suggested in [93] for testing whether all variation is selectively neutral. Bold: Significant values.