TABLE 2 in Contributions to the knowledge of the genus Trichodrilus Claparède (Lumbriculidae, Clitellata) with the description of new groundwater species from Spain

TABLE 2. Selected characters for comparison of Trichodrilus benati sp. nov., T. intermedius Hrabě and T. medius Hrabě, using the measurements made by the authors on the type material. Abbreviations: L, length; W, width; Ø, diameter; ind, individual; ant, anterior; def, vasa deferentia; post, posterior; sg, segment. Roman numbers indicate the position of the segment. Marked by—when the character could not be seen or measured CharactersT. benati sp. nov. Mairulegoreta CaveT. benati sp. nov. Azoleta CaveT. intermedius Hrabe’s collection (measured by authors)T. medius Hrabe’s collection (measured by authors)Body Ø (mm) (s. VIII to X)0.26–0.35 (0.34 whole-mount)0.35c. 0.50.45Dorsal anterior chaetae (length, µm)0.8–1 times ventrals0.8–1 times ventrals––Ventral anterior chaetae (length, µm)(70–89 in II) 96–12895 (in II) 110–132––Clitellum(post IX) X–XII (ant XIII)––X–XIIISecondary annulationFrom IIIFrom IIIFrom IIIFrom IIIPharyngeal glandsIV, V–VII (VIII ventral)IV–VIIIV–VIIII–VIPorophore/ penis (length µm)Conical/ penis 10– 17, backwardsConical/ penis 6, backwardsPenis salient c. 15Conical small/ Penis to 35Atrial ampulla (length x ø, µm)77–89 x 56–68140 x 105190 x 8294–100 x 100–102Atrial duct (µm)55–58 x 16–2480 x 3560 x 30110 x 34–67Atrial musculature thickness (µm)5–104–87–125–10Atrial epithelium (µm)12–1835–4312–1810–32Prostatic cells (height, µm)Tight clusters, to 54–73Tight clusters, to 110In clusters, to 102Tight clusters, to 135Vasa def. junctionApicalApicalApicalSubapicalVasa def. Ø µm)10–2214–1725–51 (20–22 at the junction)10–20Spermathecal ampulla (length x width, µm)90–280 x 45–74270–285 x 70–102600 x 234–261 same segment145–504 x 90 –322 May pass into adjacent segmentSpermathecal ducts (length x width, µm)40–92 x 10–21 (17–34 in vestibule)54–73 x 10–15 (26–32 in vestibule)152 x 17–48 (30 at the pore)c. 140 x 31–36SitesCave riverResurgence below a caveWells, FranceGroundwater, Germany