Natal nest locations of the Asian house rat (Rattus tanezumi) in lowland rice-coconut cropping systems: a coconut penthouse or rice bunds with water frontage?

Context. Rattus tanezumi is a serious crop pest within the island of Luzon, Philippines. In intensive flood-irrigated rice field ecosystems of Luzon, female R. tanezumi are known to primarily nest within the tillers of ripening rice fields and along the banks of irrigation canals. The nesting habits of R. tanezumi in complex rice-coconut cropping systems are unknown.