Revision of the early crocodylomorph Trialestes romeri (Archosauria, Suchia) from the lower Upper Triassic Ischigualasto Formation of Argentina: one of the oldest-known crocodylomorphs

Trialestes romeri (Reig) is an early crocodylomorph from the Ischigualasto Formation (late Carnian – early Norian; Ischigualasto – Villa Unión Basin, Argentina) and one of the oldest-known members of this clade. Two specimens of this species are known, the holotype (PVL 2561) and a referred specimen (PVL 3889), both consisting of associated cranial and postcranial remains. These specimens are incomplete and poorly preserved thus leading previous authors to propose different phylogenetic hypotheses for the species. Trialestes romeri was originally interpreted as a basal dinosaur, subsequently considered to be a crocodylomorph with some dinosaurian characters (e.g. mesotarsal ankle joint, functionally tridactyl pes), and even proposed as a chimera representing two different genera. Some recent workers have mentioned and discussed Trialestes, but none have described it in detail or included it in a quantitative phylogenetic analysis. Here, we describe in detail all the material assignable ...