Penguin virus genera and their closest relatives.

aAll viruses listed are tick borne except Rift valley fever virus. Underlined viruses or virus families are or contain, respectively, known pathogens of humans or animals.bBased on amino acid identity.cFrom ticks collected in 1972 near Royal penguin colony [3], [20].dBased on complete polyprotein sequence [4].eMay be related to Nugget and Taggert virus [3], [20], respectively.fLaboratory based experimental infection only [15].gBased on partial sequence.