TABLE 8 in Three newly discovered species of Amynthas earthworms (Oligochaeta: Megascolecidae) from Jiangxi Province and adjacent regions of China

TABLE 8. Comparative analysis of the morphological characteristics of A. shangraoensis sp. nov., A. corticis (Kinberg, 1867), A. carnosus (Goto & Hatai, 1899), A. heterochaetus (Michaelsen, 1869), and A. nanshanensis Shen, Tsai & Tsai, 2003 Morphological featureA. shangraoensis sp. nov.A. corticis (Kinberg, 1867)A. carnosus (Goto & Hatai, 1899)A. heterochaetus (Michaelsen, 1869)A. nanshanensis Shen, Tsai & Tsai, 2003Literature referencePresent studyChang et al., 2009Chang et al., 2009Xu & Xiao, 2011Chang et al., 2009Body dimensions (mm)109–147 length, 4.5–5.1 width96–184 length, 3.0–5.0 width130–240 length, 6.0–8.3 width100–158 length, 3.0–5.0 width41–89 length, 2.2–3.0 widthSecondary annulationsPresent on segments VII–XIII-Present on segments VII–XIII-Present on segments 9–13Location of first dorsal poreIntersegmental furrow 12/13Intersegmental furrows 11/12 or 12/13Intersegmental furrows 11/12 or 12/13Intersegmental furrow 11/12Intersegmental furrow 5/6Location of spermathecal pores5/6–8/95/6–8/95/6–8/95/6–8/95/6–8/9Distance between spermathecal pores0.33 × total body circumference0.28 × total body circumference0.26–0.27 × total body circumference0.57 × total body circumference0.33–0.34 × total body circumferenceGenital papillae within spermathecal pore regionA small genital papilla located anterior to the right seta b on segments VIII and IXPaired pre-setal and/or post-setal genital papillae, in line with or medial to the spermathecal poresOne genital papilla located posterior and medial to each spermathecal pore, and one or two papillae located ventral to each spermathecal poreOne genital papilla located close to the spermathecal pores and the other located on the ventral sidePresent or absent in segments VII–IXMale poreLocated in the center of a slightly raised, elliptical porophorePositioned on depressed, corneous padsPossesses a whitish spot in the center or a shallow transverse groove in the middleLarge, rounded papilla, surrounded by 3 or more concentric ridgesEach male pore is accompanied by a lateral papilla, encircled by 2–3 slight concentric foldsGenital papillae within male pore regionFlat-topped genital papillae located just posterior and medial to male pore porophorePresent or absent; if present, one or two, small, concentric papillae, located medial to male poreOne pre-setal and one post-setal genital papilla, located medial to each male pore1–2 genital papillae located near each male pore, with variable arrangementPresent or absent; located pre-setally in segments XVIII–XIX, slightly medial to the male pores, paired or single (right or left)Intestinal caecaSimpleSimpleSimpleSimpleSimpleProstate glandDeveloped, located in segments XVI to the anterior half of segment XXIWell-developed, rudimentary, or absentDeveloped, located in segments XVI– XXAlways absent, rarely present, sometimes only a vestigial structureDeveloped, located in segments XVIII–XXMorphologicalA. shangraoensis sp.A. corticisA. carnosus (GotoA. heterochaetusA. nanshanensis Shen,featurenov.(Kinberg, 1867)& Hatai, 1899)(Michaelsen, 1869)Tsai & Tsai, 2003Prostatic ductU-shaped, duct is thicker on the ventral side-U-shaped-C-shapedAccessory glands within prostate gland regionIrregular accessory gland on segment XVIIIAbsentRound, sessile, aligned with each genital papilla around the male poresRound, short ducts, associated with external papillaeSmall, stalked, associated with preclitellar papillaeSpermathecaePaired, located in segments VI–IX, progressively increasing in size from anterior to posterior segmentsPaired, located in segments VI–IX, ovoid ampullaPaired, located in segments VI–IX, 3.5–4.5 mm in length, oval ampulla with long, thick ductsPaired, located in segments VI–IX, ampulla ovoid or slender, ducts equal to or shorter than the length of the ampullaPaired, located in segments VI–IX, peach-shaped ampulla, 1.0–1.5 mm in length, well-developed ducts measuring 0.5 mm in lengthDiverticulumMeasures two-thirds to three-fourths the length of the main spermathecal axis, anterior dilation absentBlunt, ovoid, with a straight stalkMeasures three-fourths the length of the main axis, with anterior three-fifths expandedShorter than main spermathecal axis, with a round or oval seminal chamberSlender stalk, 0.5 mm in length; oval seminal chamber measuring 0.5 mmAccessory glands within spermathecal regionIrregular accessory gland in segments VIII and IXAbsentSessile/stalked, aligned with external genital papillae-Large, round, and short stalk; corresponding to external genital papillae ......continued on the next page Note: - means data not available