Observed animals during the study period in 2010.

Observed animals [sex (F = female, M = male); age in years or months (mo)] during the study period in 2010. Animals showing the throw-kiss-display are marked in capital letters; animals showing puff-blowing are marked in bold; the animal in parentheses was not observed in this study. Female Sounda is indicated to have died before the second observation period (†). Offspring are indicated underlined and in italics below their mothers. Groups 1 to 3 are the same groups as in 2000; group 5 is a new family group. Visual access was not given between the enclosures.