Coefficient estimates and adjusted Standard Errors (SE) of environmental variables (forest area, AWMSI, temperature, rainfall and TWI) after model averaging of GLMMs to explore their relationship with tree, millipede and bird species richness.

The response variables in all models were detrended species richness (i.e. residuals from spatial and non-spatial models). Estimates represent standardize values and, therefore, they are in a comparable scale. The relative importance of predictor variables was calculated as the sum of AICc-wi values of the competing models in which the predictor was present. Significant predictors at P