Research applications addressed by dispersal-related papers*, the number of papers addressing each in the Old and New time periods.

P = P value of Fisher's Exact test. There were also no significant differences when questions addressed by less than 10 papers were excluded from the test. Note there were only 18 completed reviews among old land planning papers so old and new numbers are not directly comparable.*Our categories of research were developed during workshops with the authors and were based on one-third of the papers that we reviewed. The research questions that we identified therefore arise from the papers. However, we acknowledge that there may be other ways that the broad categories of research might be defined, and this may reveal different insights into the nature of changing research questions over time.