Beetles, barcodes, and big data: a deep dive into the phylogeny of Harpalinae (Carabidae)

The ground beetles (Carabidae) are a highly species-rich lineage of the Coleoptera, with over half of their diversity concentrated in the ~20,000 described species in the subfamily Harpalinae sensu lato. As a presumed recent radiation lacking deeply distinct morphological divisions, their taxonomic classification has been challenging, while molecular studies remain limited in the number of genes and taxa sampled. Using ~450 mitochondrial genome sequences from across the Carabidae and the major biogeographic realms we investigate the tribal relationships in Harpalinae. Our phylogenetic analysis supports a revised system that broadly divides the harpalines into two major reciprocally monophyletic lineages, corresponding to a narrowly defined Harpalinae sensu novo and a distinct Lebiinae. Within Harpalinae, we recover well-supported subclades that mostly represent existing tribes (e.g., Harpalini, Pterostichini, Licinini, Platynini), while clades in Lebiinae required the recognition of thr..., , # Beetles, barcodes, and big data: a deep dive into the phylogeny of Harpalinae (Carabidae)

Authors: Beulah Garner, Aileen Scott and Alfried Vogler

Journal: Systematic Entomology

Dataset DOI: [10.5061/dryad.tht76hfdk](https://doi.org/10.5061/dryad.tht76hfdk)

This dataset contains multiple-sequence alignments, partitions schemes, phylogenies and metadata from the above paper.

## archive_scripts_carabidae_phylogeny_260302.zip

An archive of all original scripts used for this paper. See README file within for more detail. Also available at [https://github.com/beetlephylo/carabidae_phylogeny](https://github.com/beetlephylo/carabidae_phylogeny)

## File_R1_raw_sequences_2024.zip

The unprocessed sequence files resulting from the BLAST and BOLD searches carried out in 2024. These sequences were processed and used for the mitogenome phylogeny as described in File_S1_methods.docx

## File_R2_raw_sequences_2025.zip

The unprocessed seqeunce files resulting from the GenBank and BOLD searches..., ,