Macroinvertebrates indicated by BVSTEP as associated with gradients of specific predictor variables in autumn and spring.

ρ signifies Spearman’s correlations and P-value shows the significance of the relationship between the macroinvertebrates and the predictor variables. Blank cells between taxon and predictor variable indicate that taxon was not correlated with that predictor variable.Numbers written in the cells are Spearman’s correlation values between the taxon and gradient of that predictor variable. Predictor variables for both seasons are arranged in the order in which the most influential variables in each season appears as indicated by DistLM (Table 4) appear. Abbreviations for predictor variables are listed in S1 Table.