Density-dependent feedback across nested levels of organization in a social primate

The strength and direction of density-dependent mechanisms acting on individual reproduction and survival may vary across the nested levels of organization social animals live in, such that complex patterns of density dependence shape fitness and population growth. Yet knowledge of such processes of population regulation where individuals are simultaneously subjected to contrasting density effects remains limited. We quantify and contrast density effects on components of individual fitness across two nested levels of organization: the population and the social group, using 45 years of demographic data of rhesus macaques. Our analysis reveals opposing density feedback on individual reproduction and survival across levels of organization and shows that density does not affect all life stages equally. While increased population density reduced female reproduction during maturation, females in larger groups were more likely to reproduce. Infant survival was optimal at intermediate populatio..., , , # Data from: Density-dependent feedback across nested levels of organization in a social primate

Data and codes provide examples for estimating the main results presented in tables and figures.

## Description of the data and file structure

Code files:
Reproduction_Analysis.R - These codes provide the workflow for estimating population- and group-level density-dependent feedback on individual reproduction using GLMMs.
Survival_Analysis.R - These codes provide the workflow for estimating population- and group-level density-dependent feedback on individual survival using GLMMs.

### Data files:

Reproduction_Data.RData - This data frame contains annual individual reproduction status and adult population and adult group density during the 44 year study. year column represents the year the data was recorded in. id represents the ID of each monkey (deidentified). fertility equals 1 if female produced offspring; 0 = if female did not produce offspring. mom represents the ID of the monkey's ...,