Parameter structure for multistate models fit to the data for this study.

Ages represent separation of years with older ages than those listed being grouped as similar. Ss and Sw represent structure for survival of those that were suckling and those that were weaned in year i – 1, respectively. Sighting probability structure for individuals that were weaned (pw) is represented by p and those that were suckling (ps) constrained to 1. State transitions from suckling to independence (ψsw) are represented by ψ, while the reverse (ψws) was constrained to 0.