TABLE 1 in Genus-level revision of the Heterozerconoidea (Parasitiformes: Mesostigmata)

TABLE 1. Characters and character states in phylogenetic analysis of relationships in Heterozerconoidea.
Char. #Character description and statesGNATHOSOMA1Basal part of fixed digit in female: 0, stout, length width ratio 1–2 (Figs. 1, 3); 1, elongate, length width ratio 4–6 (Fig. 2).2Movable digit in adults: 0, without thin, strap-like extensions; 1, with thin, strap-like extensions (Figs. 1–3, red arrows).3Movable digit in female: 0, stout, digit length ~3 times basal width (Fig. 1); 1, elongate, digit length>6 times basal width (Figs. 2–3).4Inside movable digit female: 0, with distinct teeth; 1, with brush-like structure (Fig. 3, blue arrow).5Excrescences on male chelicera: 0, absent; 1, present, on movable digit; 2, present, interdigital.6Spermatodactyl on chelicera male: 0, absent; 1, present, on movable digit; 2, present, on fixed digit.7Spermatodactyl shape: 0, straight (Figs. 4–7, 16–19); 1, flaccid (Figs. 8–9, 14–15); 2, recurved (Figs. 10–11); 3, compressed (Figs. 12–13).8Axial outgrowth of palp trochanter in female: 0, absent; 1, small and membranous; 2, distinct and membranous (e.g., Figs. 22, 37); 3, distinct and sclerotized (Fig. 29, red arrow).9Axial outgrowth of palp trochanter in male: 0, absent; 1, small and membranous; 2, distinct and membranous (e.g., Fig. 42); 3, distinct and sclerotized (Fig. 40).10Seta pd1 on palp femur: 0, absent; 1, present.11Setae on palp femur: 0, relatively thin, setiform (Fig. 21); 1, thick, spine-like (Fig. 22).12Seta al2 on palp genu: 0, present; 1, absent.13Seta pl on palp genu: 0, present; 1, absent (Fig. 22).14Number of sensilla on palp tarsus: 0, 13–15; 1, 10–12; 2, 9.15Gnathotectum: 0, without prominent points; 1, with prominent points.16Anterior margin gnathotectum: 0, serrate 1, not serrate.17Overall shape of gnathotectum: 0, triangular; 1, curved; 2, blunt.18Sexual differentiation in gnathotectum shape: 0, absent; 1, present.19Lateral lips: 0, independent small flaps; 1, enlarged to form a trough (Figs. 23–42).20Posterior extension of lateral lips: 0, short, not extending beyond insertions of setae hyp2 (Fig. 30); 1, long, extending distinctly past insertions of setae hyp 2 (Fig. 32).21Distance from setae hyp3 to sc vs. hyp3 to hyp2: 0, distinctly smaller; 1, subequal; 2, distinctly larger.22Length setae hyp2 relative to hyp 3: 0, subequal or slightly longer; 1, at least twice as long.23Cornicula: 0, solid; 1, membranous.24Membranous cornicula: 0, without a distal notch; 1, with a distal notch (Fig. 34, arrow).25Additional setal base-like structure on hypostome: 0, absent; 1, present (Fig. 35, arrow).IDIOSOMA26Pygidial shield in larva: 0, present; 1, absent.27Pygidial shield in protonymph: 0, present; 1, absent.28Anterior dorsal setae j1 (Figs. 45–46): 0, long (> 35 µm); 1, medium long (10–30 µm); 2, minute (<5 µm)29Anterior dorsal margin in female: 0, with at most a single pair of elongate setae (j1); 1, with multiple medium to long setae in j1 position (Fig. 48).30Antero-lateral margin of dorsal shield in males. 0, without distinct spines; 1, with small, hook-like spines (Fig. 49).31Median dorsal setae. 0, distinct; 1, minute32Posterior dorsal setae Z2–Z4. 0, elongate (Figs. 45–46); 1, minute (Fig. 47).33Areas near sternal seta st1 in female. 0, sclerotized; 1, with soft cuticle.66Sternitiventral and anal shields in male: 0, not fused; 1, fused, but line of fusion still visible; 2, fused, line of fusion not visible.67Setae Sv2 in female: 0, inserted on genitiventral shield; 1, not inserted on genitiventral shield.68Setae Sv2 in male: 0, inserted on sternitiventral shield; 1, not inserted on sternitiventral shield.69Setae Sv3 in female: 0, inserted on genitiventral shield; 1, not inserted on genitiventral shield70Setae Sv3 in male: 0, inserted on sternitiventral shield; 1, not inserted on sternitiventral shield71Insertion of setae Jv5: 0, anterior to ventrianal fusion line (Fig. 57); 1, posterior to ventrianal fusion line (Fig. 69).72Insertion paranal setae (pa): 0, at level of anus; 1, posterior to the anus; 2, anterior to the anus.73Postanal (po) seta: 0, of similar length as paranal (pa) setae; 1, less than 1/3 the length of paranal (pa) setae.74Setae Z5 in male: 0, elongate (>40 µm); 1, medium but distinct (15–40 µm); 2, small to minute (<15 µm).75Setae S5 in male: 0, elongate (>40 µm); 1, medium but distinct (15–40 µm); 2, small to minute (<15 µm).76Equally spaced long marginal setae: 0, absent; 1, present (Fig. 43).77Marginal opisthosomal setae: 0, all setiform; 1, some spine-like; 2, some flattened (Figs. 43, 53–54).78Shape of spine-like marginal opisthosomal setae: 0, peg-like (Figs. 59–60); 1, hook-like (Figs. 65–66); 2, anchor-like (Figs. 63–64). Inapplicable if setae not spine-like.79Lyrifissures iv5 in male: 0, inserted near anterior margin of sucker; 1, absent.80Postero-marginal shields in female: 0, small, not extending lateral to opisthogastral suckers (Fig. 57); 1, wide, extending lateral well beyond opisthogastral suckers.LEGS81Tibiae and tarsi of legs I: 0, of similar width as the rest of the leg; 1, somewhat narrowed relative to the rest of the leg; 2, distinctly narrowed relative to the rest of the leg (Fig. 85).82Acrotarsus on legs I: 0, present (Fig. 73); 1, absent.83Femora I seta ad3: 0, absent; 1, present.84Femora I setae v3, v4: 0, present; 1, absent.85Femora I seta pl2: 0, absent; 1, present.86Genua I seta ad3, pd3: 0, present; 1, absent.87Tibiae I seta ad2: 0, present; 1, absent.88Tibiae I seta ad3, pd3: 0, present; 1, absent.89Tibiae I setae av2, pv2: 0, present; 1, absent.90Femora II seta av2: 0, present; 1, absent.91Femora II seta pv2: 0, present; 1, absent.92Genua II–III seta ad3: 0, present; 1, absent.93Genua II seta pd3: 0, present; 1, absent.94Genua II seta pl2: 0, present; 1, absent.95Tibiae II–IV seta ad2: 0, present; 1, absent..96Tibiae II–IV seta pd3: 0, present; 1, absent.97Tibiae II–IV seta pl2: 0, present; 1, absent.98Femora III seta v3: 0, present; 1, absent.99Genua IV seta a d3: 0, present; 1, absent.100Genua III–IV seta pd3: 0, present; 1, absent.101Tarsi IV setae av4, pv4: 0, present; 1, absent.102Coxae I setae in male. 0, setiform; 1, distinct spines (Figs. 67–68).103Femora I setae al1 and al2 in male. 0, setiform, in anterolateral position; 1, spinose, in anteroventral position (Fig. 84).104Femora I seta av in male. 0, setiform; 1, large, curved spine (Figs. 73, 79); 2, large straight spine (Figs. 80– 86).105Femora II seta al in male. 0, setiform; 1, spine and ventral in position (Figs. 93–96).106Femora II seta pl in male. 0, setiform; 1, spine and ventral in position (Figs. 93–96).107Femora II seta av in male. 0, setiform; 1, spine (Figs. 93–96).108Femora II seta pv in male. 0, setiform; 1, spine (Figs. 93–96).109Genua II seta pv in male. 0, setiform; 1, spine (Figs. 94–95).110Tibiae II seta pv in male. 0, setiform; 1, spine (Figs. 94–95).111Tarsi II seta pl1. 0, setiform; 1, spine (Figs. 89–90, 93–94).112Trochanters III seta al in male. 0, setiform; 1, distinct spine.113Trochanters III seta av in male. 0, setiform; 1, distinct spine.114Trochanters III seta pl in male. 0, setiform; 1, short, thick, rounded spine; 2, distinct spine, with pointed tip.
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