Table 4 in Systematics of Huicundomantis, a new subgenus of Pristimantis (Anura, Strabomantidae) with extraordinary cryptic diversity and eleven new species

Table 4. Qualitative morphological traits of Pristimantis, subgenus Huicundomantis. Coloration corresponds to live individuals unless otherwise noticed. + is for present; - is for absent. Dorsal textureCranial crestsPostocular/scapular fold/ridgeMiddorsal foldTubercles row headDorsolateral foldsLateral foldSnout shape (dorsal; lateral)Discs (expansion, shape)Basal webbingIris colorationGroin colorationP. atillo sp. nov.Shagreen with or without scattered small subconical tubercles--+/-+-+/-Acuminate; protruding; with a fleshy keelExpanded to broadly expanded, rounded to elliptical+Copper with a faint medial horizontal darker streakBright orange, surrounded or not by yellow spotsP. atratusShagreenLow-+/-+/-++Subacuminate; rounded; with or without a papillaBroadly expanded, rounded to elliptical+Pale yellow to pale bronze, thin brown streakBlack with white or yellow spotsP. balionotusTuberculate-----+/-Subacuminate; rounded; with a papillaSlightly expanded, elliptical-Bronze with a red medial streakIn preservative, pale rusty brown with or without faint brown marblingP. chomskyi sp. nov.Shagreen------Subacuminate; rounded; with or without a papillaExpanded, rounded to elliptical+Orange with a faint reddish brown streakDark chocolate brown with or without small cream flecksP. gloria sp. nov.Tuberculate to warty--+---Subacuminate; rounded; with or without a papillaExpanded, truncate to elliptical+Light silver to cream with wide black reticulations and a red streakPinkish to purplish brown with irregular cream to light brown flecks or spotsP. hampatusamiShagreen with scattered tubercles-Low, W-shaped++-+Subacuminate; rounded; with or without a papillaBroadly expanded, elliptical to truncate-Golden to bronze with a red medial horizontal streakReddish brown with yellow spotsP. jimenezi sp. nov.Shagreen with or without scattered small subconical tubercles--++-+Acuminate; protruding; with a fleshy keelExpanded to broadly expanded, elliptical to truncate+Copper with red medial streak to completely redPinkish, purplish or dark brown with small light brown to yellow spotsP. lutzae sp. nov.Shagreen to tuberculate--+/-+/--+/-Round to subacuminate, with or without a papillaExpanded, elliptical to truncate+Golden to creamy brown with a reddish brown medial streakPinkish to reddish brown, suffused or not with orange, with cream or light brown spotsP. multicolor sp. nov.Shagreen to warty---+/---Subacuminate; rounded; with or without a papillaExpanded to broadly expanded, elliptical to truncate+Copper to creamy yellow with or without a red to dark brown streakCream, orange, brown, or black with or without cream to yellow flecks or spotsP. phoxocephalusShagreen with scattered small tubercles--++-+/-Acuminate; protruding; with a fleshy keelBroadly expanded, rounded to elliptical+Golden with wide black reticulationsIn preservative, brown with cream reticulation or spotsP. percultusTuberculateLow-----Subacuminate; acutely rounded; with a keelExpanded, elliptical+Copper with or without a faint red medial streakYellow with black reticulationsP. teslai sp. nov.Tuberculate with prominent and rounded tubercles---+--Acuminate; protruding; with a fleshy keelExpanded to broadly expanded, elliptical to truncate+CopperDark brown with yellow irregular blotchesP. tinguichacaSmooth--++-+Subacuminate; rounded; with or without a papillaBroadly expanded, rounded+Red with a dark medial streakReddish to dark brownP. torresi sp. nov.Shagreen--++/--+Acuminate; protruding; with a fleshy keelBroadly expanded, elliptical to truncate+Golden to beige with red to reddish brown streakLight purplish brown to brown with or without yellow spotsP. totoroi sp. nov.Shagreen with or without scattered small tubercles--++-+Acuminate; protruding; with a fleshy keelBroadly expanded, rounded to elliptical+Golden with a medial horizontal red streakIn preservative, brown with or without pale spotsP. verrucolatus sp. nov.Shagreen with or without scattered small tubercles--+/-+/--+Acuminate; rounded; with a fleshy keelBroadly expanded, elliptical to truncate+Coppery brownReddish brown with small light brown, orangey brown or yellow spotsP. versicolorShagreen to tuberculate--+/----Subacuminate; rounded; with or without a papillaExpanded, rounded to elliptical+Bronze-white with a reddish brown streakBrown to black with cream to reddish pink flecks or spotsP. philipiTuberculate with low tubercles and warts-W-shaped ridge+/-+--RoundedSlightly expanded, truncate-Grayish bronzeBlack with or without white or yellow streaksP. cryptomelasShagreen-Prominent,) (-shaped+/-+-+Subacuminate; rounded; with or without a papillaBroadly expanded, rounded to elliptical+Red, orange or cream with a red or orange streakBlack with or without yellow or white spotsP. gagliardoiShagreen with scattered tubercles-Prominent, W-shaped+/-+-+RoundedBroadly expanded, elliptical to truncate+Bronze with or without a brown medial horizontal streakPink to orange with or without brown blotchesP. muscosusSmooth to shagreen-Low,) (-shaped-+-+RoundedBroadly expanded, elliptical to truncate+Reddish brownDark brown with orange, yellow or white spotsP. nangaritza sp. nov.Finely tuberculate-Low,) (-shaped+/-+-+/-Subacuminate; roundedBroadly expanded, rounded to elliptical+UnknownIn preservative, light brown with or without pale flecksP. spinosusFinely tuberculateLowLow,) (-shaped-+-+Subacuminate; rounded to truncateBroadly expanded, rounded+UnknownBlack with big white spots表4 惠昆蛙亚属（Huicundomantis）普氏蛙属（Pristimantis）的定性形态性状。除特别注明外，体色均为活体个体的真实色泽。符号"+"表示该性状存在，"-"表示该性状不存在。本次记录的性状依次为：背部纹理、颅顶嵴、眼后/肩褶/脊、背中褶、头部结节列、背侧褶、侧褶、吻部形态（背面观；侧面观）、指（趾）盘（扩张程度、形态）、基部蹼、虹膜色泽、股部内侧色泽。 1. P. atillo 新种（sp. nov.）：背部纹理为具沙粒状质感，或散布少量亚圆锥状结节；颅顶嵴缺失；眼后/肩褶/脊缺失；背中褶存在/缺失；头部结节列存在；背侧褶缺失；侧褶存在/缺失；吻部呈渐尖形，突出，具肉质脊；指（趾）盘扩张至明显扩张，呈圆形至椭圆形；基部蹼存在；虹膜呈铜色，具微弱的中部水平深色条纹；股部内侧为亮橙色，周围有或无黄色斑点。 2. P. atratus：背部纹理为具沙粒状质感；颅顶嵴低平；眼后/肩褶/脊存在/缺失；背中褶存在/缺失；头部结节列存在；背侧褶存在；侧褶存在；吻部近渐尖形，圆形，具或不具乳突；指（趾）盘明显扩张，呈圆形至椭圆形；基部蹼存在；虹膜呈淡黄色至淡铜色，具细褐色条纹；股部内侧为黑色，具白色或黄色斑点。 3. P. balionotus：背部纹理为具结节状突起；颅顶嵴缺失；眼后/肩褶/脊缺失；背中褶缺失；头部结节列缺失；背侧褶缺失；侧褶存在/缺失；吻部近渐尖形，圆形，具乳突；指（趾）盘轻微扩张，呈椭圆形；基部蹼缺失；虹膜呈铜色，具红色中部条纹；经防腐固定后呈淡锈褐色，具或不具模糊的褐色大理石纹。 4. P. chomskyi 新种（sp. nov.）：背部纹理为具沙粒状质感；颅顶嵴缺失；眼后/肩褶/脊缺失；背中褶缺失；头部结节列缺失；背侧褶缺失；侧褶缺失；吻部近渐尖形，圆形，具或不具乳突；指（趾）盘扩张，呈圆形至椭圆形；基部蹼存在；虹膜呈橙色，具微弱的红棕色条纹；股部内侧为深巧克力褐色，具或不具细小奶油色斑点。 5. P. gloria 新种（sp. nov.）：背部纹理为具结节至具疣状突起；颅顶嵴缺失；眼后/肩褶/脊缺失；背中褶存在/缺失；头部结节列缺失；背侧褶缺失；侧褶缺失；吻部近渐尖形，圆形，具或不具乳突；指（趾）盘扩张，呈截形至椭圆形；基部蹼存在；虹膜呈浅银色至奶油色，具宽阔的黑色网状纹及红色条纹；股部内侧呈粉红色至紫褐色，具不规则奶油色至浅褐色斑点。 6. P. hampatusami：背部纹理为具沙粒状质感且散布结节；颅顶嵴缺失；眼后/肩褶/脊低平，呈W形；背中褶存在；头部结节列存在；背侧褶缺失；侧褶存在；吻部近渐尖形，圆形，具或不具乳突；指（趾）盘明显扩张，呈椭圆形至截形；基部蹼缺失；虹膜呈金色至铜色，具红色中部水平条纹；股部内侧为红棕色，具黄色斑点。 7. P. jimenezi 新种（sp. nov.）：背部纹理为具沙粒状质感，或散布少量亚圆锥状结节；颅顶嵴缺失；眼后/肩褶/脊缺失；背中褶存在；头部结节列存在；背侧褶缺失；侧褶存在；吻部呈渐尖形，突出，具肉质脊；指（趾）盘扩张至明显扩张，呈椭圆形至截形；基部蹼存在；虹膜呈铜色，具红色中部条纹或通体红色；股部内侧呈粉红色、紫褐色或深褐色，具细小浅褐色至黄色斑点。 8. P. lutzae 新种（sp. nov.）：背部纹理为具沙粒状质感至具结节；颅顶嵴缺失；眼后/肩褶/脊存在/缺失；头部结节列存在/缺失；背侧褶缺失；侧褶缺失；吻部呈圆形至近渐尖形，具或不具乳突；指（趾）盘扩张，呈椭圆形至截形；基部蹼存在；虹膜呈金色至奶油褐色，具红棕色中部条纹；股部内侧呈粉红色至红棕色，具或不具橙色浸染，具奶油色或浅褐色斑点。 9. P. multicolor 新种（sp. nov.）：背部纹理为具沙粒状质感至具疣状突起；颅顶嵴缺失；眼后/肩褶/脊缺失；背中褶缺失；头部结节列存在/缺失；背侧褶缺失；侧褶缺失；吻部近渐尖形，圆形，具或不具乳突；指（趾）盘扩张至明显扩张，呈椭圆形至截形；基部蹼存在；虹膜呈铜色至奶油黄色，具或不具红色至深褐色条纹；股部内侧呈奶油色、橙色、褐色或黑色，具或不具奶油色至黄色斑点。 10. P. phoxocephalus：背部纹理为具沙粒状质感且散布细小结节；颅顶嵴缺失；眼后/肩褶/脊缺失；背中褶存在；头部结节列存在；背侧褶缺失；侧褶存在/缺失；吻部呈渐尖形，突出，具肉质脊；指（趾）盘明显扩张，呈圆形至椭圆形；基部蹼存在；虹膜呈金色，具宽阔的黑色网状纹；经防腐固定后呈褐色，具奶油色网状纹或斑点。 11. P. percultus：背部纹理为具结节状突起；颅顶嵴低平；眼后/肩褶/脊缺失；背中褶缺失；头部结节列缺失；背侧褶缺失；侧褶缺失；吻部近渐尖形，急尖，具脊；指（趾）盘扩张，呈椭圆形；基部蹼存在；虹膜呈铜色，具或不具微弱的红色中部条纹；股部内侧呈黄色，具黑色网状纹。 12. P. teslai 新种（sp. nov.）：背部纹理为具明显且圆形的结节状突起；颅顶嵴缺失；眼后/肩褶/脊缺失；背中褶缺失；头部结节列存在；背侧褶缺失；侧褶缺失；吻部呈渐尖形，突出，具肉质脊；指（趾）盘扩张至明显扩张，呈椭圆形至截形；基部蹼存在；虹膜呈铜色；股部内侧为深褐色，具黄色不规则斑块。 13. P. tinguichaca：背部纹理为光滑；颅顶嵴缺失；眼后/肩褶/脊缺失；背中褶存在；头部结节列存在；背侧褶缺失；侧褶存在；吻部近渐尖形，圆形，具或不具乳突；指（趾）盘明显扩张，呈圆形；基部蹼存在；虹膜呈红色，具深色中部条纹；股部内侧呈红色至深褐色。 14. P. torresi 新种（sp. nov.）：背部纹理为具沙粒状质感；颅顶嵴缺失；眼后/肩褶/脊缺失；背中褶存在/缺失；头部结节列缺失；背侧褶缺失；侧褶存在；吻部呈渐尖形，突出，具肉质脊；指（趾）盘明显扩张，呈椭圆形至截形；基部蹼存在；虹膜呈金色至米黄色，具红色至红棕色条纹；股部内侧呈浅紫褐色至褐色，具或不具黄色斑点。 15. P. totoroi 新种（sp. nov.）：背部纹理为具沙粒状质感，或散布少量结节；颅顶嵴缺失；眼后/肩褶/脊缺失；背中褶存在；头部结节列存在；背侧褶缺失；侧褶存在；吻部呈渐尖形，突出，具肉质脊；指（趾）盘明显扩张，呈圆形至椭圆形；基部蹼存在；虹膜呈金色，具中部水平红色条纹；经防腐固定后呈褐色，具或不具浅色斑点。 16. P. verrucolatus 新种（sp. nov.）：背部纹理为具沙粒状质感，或散布少量结节；颅顶嵴缺失；眼后/肩褶/脊存在/缺失；头部结节列存在/缺失；背侧褶缺失；侧褶缺失；吻部呈渐尖形，圆形，具肉质脊；指（趾）盘明显扩张，呈椭圆形至截形；基部蹼存在；虹膜呈铜褐色；股部内侧呈红棕色，具细小浅褐色、橙棕色或黄色斑点。 17. P. versicolor：背部纹理为具沙粒状质感至具结节；颅顶嵴缺失；眼后/肩褶/脊存在/缺失；背中褶缺失；头部结节列缺失；背侧褶缺失；侧褶缺失；吻部近渐尖形，圆形，具或不具乳突；指（趾）盘扩张，呈圆形至椭圆形；基部蹼存在；虹膜呈铜白色，具红棕色条纹；股部内侧呈褐色至黑色，具奶油色至浅红色斑点。 18. P. philipi：背部纹理为具低结节及疣状突起；颅顶嵴缺失；眼后/肩褶/脊呈W形；背中褶存在/缺失；头部结节列存在；背侧褶缺失；侧褶缺失；吻部呈圆形；指（趾）盘轻微扩张，呈截形；基部蹼缺失；虹膜呈灰铜色；股部内侧为黑色，具或不具白色或黄色条纹。 19. P. cryptomelas：背部纹理为具沙粒状质感；颅顶嵴明显，呈(-)形；眼后/肩褶/脊存在/缺失；背中褶存在；头部结节列缺失；侧褶存在；吻部近渐尖形，圆形，具或不具乳突；指（趾）盘明显扩张，呈圆形至椭圆形；基部蹼存在；虹膜呈红色、橙色或奶油色，具红色或橙色条纹；股部内侧为黑色，具或不具黄色或白色斑点。 20. P. gagliardoi：背部纹理为具沙粒状质感且散布结节；颅顶嵴明显，呈W形；眼后/肩褶/脊存在/缺失；背中褶存在；头部结节列缺失；侧褶存在；吻部呈圆形；指（趾）盘明显扩张，呈椭圆形至截形；基部蹼存在；虹膜呈铜色，具或不具褐色中部水平条纹；股部内侧呈粉色至橙色，具或不具褐色斑块。 21. P. muscosus：背部纹理为光滑至具沙粒状质感；颅顶嵴低平，呈(-)形；眼后/肩褶/脊缺失；背中褶存在；头部结节列缺失；侧褶存在；吻部呈圆形；指（趾）盘明显扩张，呈椭圆形至截形；基部蹼存在；虹膜呈红棕色；经防腐固定后呈深褐色，具橙色、黄色或白色斑点。 22. P. nangaritza 新种（sp. nov.）：背部纹理为具细结节；颅顶嵴低平，呈(-)形；眼后/肩褶/脊存在/缺失；背中褶存在/缺失；头部结节列缺失；侧褶存在/缺失；吻部近渐尖形，圆形；指（趾）盘明显扩张，呈圆形至椭圆形；基部蹼存在；虹膜色泽未知；经防腐固定后呈浅褐色，具或不具浅色斑点。 23. P. spinosus：背部纹理为具细结节；颅顶嵴低平；眼后/肩褶/脊呈(-)形；背中褶缺失；头部结节列存在；侧褶存在；吻部近渐尖形，圆形至截形；指（趾）盘明显扩张，呈圆形；基部蹼缺失；虹膜色泽未知；股部内侧为黑色，具大型白色斑点。