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TABLE 1 in A new species of Helietta (Pilocarpinae, Zanthoxyloideae, Rutaceae) from Colombia and notes on the morphology and circumscription of H. glaziovii

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Zenodo2025-04-05 更新2026-05-26 收录
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TABLE 1. Comparative characters of Helietta apiculata, H. glaziovii, and H. magna. SpeciesCharacterH. apiculataH. glazioviiH. magnaLength of leaflets (cm)(3–)5–8.5(–9.5)3–8(–9)(4–)5–12(–17.3)Width of leaflets (cm)0.5–1.7(–2.5)1.4–3.4(–4.2)(1.1–)2.1–5.6(–7)Apex of leafletsApiculate with drip tip, margin involute along entire length of apiculum and adjacent marginsRounded to blunt or retuse to emarginate, margin involute laterally at very apex or not involuteAcuminate without drip tip, margin involute only over the acumenMerosity of flowers(4–)5-merous(4–)5-merous5-merousIndumentum of petalsGlabrousGlabrous or ciliolateGlabrousIndumentum of ovaryGlabrousGlabrousPuberulous apicallyShape of wing of mericarpsOblongOblongObovate to ellipticWidth of wing of mericarps (mm)5–99–11(8–)10–13DistributionEastern Paraguay, southern Brazil and in northeastern Argentina*Central and eastern BrazilCentral Colombia *The previous record of Helietta apiculata from Peru is excluded here. This record was based on Gentry & Smith 45084 (MO-n.v., U- 1578709 [digital image!]), which only have young inflorescences and the leaf morphology does not completely match H. apiculata. In addition, leaflets with involute margin at the apex have been detected in some species of Esenbeckia Kunth (1825: 246), e.g. in E. cornuta Engler (1874: 146), and occasionally in Balfourodendron Corr. Méllo ex Oliver (in Hooker 1877: 3–4). It is considered here that assign a precise identity to the specimen Gentry & Smith 45084 is not possible while fruiting specimens from that population have not seen.

表1. 尖叶赫列塔(Helietta apiculata、H. glaziovii与大赫列塔(H. magna)的比较形态特征。 | 特征项 | 尖叶赫列塔(Helietta apiculata) | 格氏赫列塔(H. glaziovii) | 大赫列塔(H. magna) | | --- | --- | --- | --- | | 小叶长度(cm) | (3–)5–8.5(–9.5) | 3–8(–9) | (4–)5–12(–17.3) | | 小叶宽度(cm) | 0.5–1.7(–2.5) | 1.4–3.4(–4.2) | (1.1–)2.1–5.6(–7) | | 小叶先端 | 具短尖并带有滴水尖,沿短突全长及相邻叶缘内卷 | 先端圆形至钝形,或微凹至凹缺,仅在极先端两侧叶缘内卷或不外卷 | 渐尖,无滴水尖,仅在渐尖部分叶缘内卷 | | 花基数 | (4–)5基数 | (4–)5基数 | 5基数 | | 花瓣被毛情况 | 无毛 | 无毛或具缘毛 | 无毛 | | 子房被毛情况 | 无毛 | 无毛 | 先端被微柔毛 | | 分果翅形状 | 长圆形 | 长圆形 | 倒卵形至椭圆形 | | 分果翅宽度(mm) | 5–9 | 9–11 | (8–)10–13 | | 分布区 | 巴拉圭东部、巴西南部及阿根廷东北部* | 巴西中部及东部 | 哥伦比亚中部 | *此前产自秘鲁的尖叶赫列塔(Helietta apiculata)记录已被排除在本次统计之外。该记录依据Gentry & Smith 45084号标本(MO馆藏未标注,U-1578709 [数字影像!],该标本仅带有幼花序,且叶片形态与尖叶赫列塔的特征不完全吻合。此外,部分埃森贝克木属(Esenbeckia Kunth, 1825: 246)物种(如角状埃森贝克木(E. cornuta Engler, 1874: 146))以及偶尔鲍弗罗登德龙属(Balfourodendron Corr. Méllo ex Oliver, in Hooker 1877: 3–4)的物种,其小叶先端叶缘内卷的情况。本文认为,因尚未采集到该种群的果期标本,因此无法对Gentry & Smith 45084号标本作出精确的物种鉴定。

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