Table 5 in An investigation into the taxonomy of Abavorana luctuosa (Peters, 1871) (Anura, Ranidae) and the resurrection of Rana decorata Mocquard, 1890 from Borneo

Table 5. Summary statistics and ventral colour pattern among members of the genus Abavorana. Summary statistics and ventral colour pattern for Abavorana decorata, A. nazgul, and A. luctuosa populations from the Malay Peninsula and Borneo, SD = standard deviation, N = sample size, / = no data. Humeral gland only present in males and all measurements in mm. A. decorataA. nazgulA. luctuosa Malay PeninsulaA. luctuosa BorneoSexMaleFemaleMaleFemaleMaleFemaleMaleFemaleN267110112211Snout-vent length (SVL)mean (± SD)51.11 (± 6.92)48.73 (± 13.32)42.77 (± 1.22)52.5140.74 (± 2.54)47.38 (± 2.36)38.15 (± 1.45)43.23 (± 2.51)range46.21-56.0029.66-58.4241.51-44.10/37.20-44.2743.89-51.0635.05-40.5537.05-45.20Head length (HL)mean (± SD)19.10 (± 0.06)18.95 (± 4.85)16.61 (± 0.25)19.6515.64 (± 0.81)17.72 (± 1.39)15.25 (± 0.78)16.88 (± 0.92)range19.05-19.1412.23-23.7116.36-17.14/14.45-16.6915.20-19.5913.53-16.7814.84-17.72Head width (HW)mean (± SD)18.60 (± 1.32)17.18 (± 4.68)15.16 (± 0.46)18.5714.24 (± 0.66)15.77 (± 1.01)13.39 (± 0.52)14.69 (± 0.83)range17.66-19.5310.62-21.0714.63-15.83/13.35-15.5813.83-16.8412.25-14.4112.71-15.74Snout length (SL)mean (± SD)7.99 (± 0.33)7.73 (± 1.85)7.31 (± 0.20)8.696.88 (± 0.43)7.82 (± 0.56)6.55 (± 0.34)7.17 (± 0.33)range7.76-8.224.69-9.467.10-7.57/6.20-7.566.48-8.305.66-7.036.61-7.86Snout width (SW)mean (± SD)10.31 (± 0.57)9.54 (± 2.28)8.26 (± 0.30)9.598.08 (± 0.44)8.84 (± 0.40)7.73 (± 0.23)8.33 (± 0.51)range9.91-10.716.25-11.687.98-8.70/7.11-8.628.21-9.387.33-8.227.65-9.02Inter orbital distance (IOD)mean (± SD)5.67 (± 0.40)5.63 (± 1.22)4.20 (± 0.23)4.944.35 (± 0.52)4.93 (± 0.54)4.24 (± 0.30)4.69 (± 0.37)range5.59-5.953.91-6.783.94-4.48/3.25-5.143.80-5.643.78-4.704.15-5.60Internarial distance (IND)mean (± SD)6.13 (± 0.05)5.76 (± 1.34)5.64 (± 0.13)6.615.33 (± 0.30)6.06 (± 0.38)5.16 (± 0.24)5.50 (± 0.42)range6.09-6.163.68-6.775.45-5.80/4.93-5.885.42-6.724.66-5.584.75-5.99Eye diameter (ED)mean (± SD)5.75 (± 0.26)5.31 (± 1.10)4.17 (± 0.24)4.804.25 (± 0.26)4.87 (± 0.29)4.18 (± 0.26)4.69 (± 0.29)range5.56-5.933.94-6.583.86-4.56/3.80-4.704.38-5.213.65-4.594.08-5.22Tympanum diameter (TD)mean (± SD)4.57 (± 0.04)4.30 (± 1.43)3.72 (± 0.19)4.103.30 (± 0.31)3.87 (± 0.27)3.16 (± 0.29)3.44 (± 0.25)range4.54-4.602.27-5.633.52-4.01/2.62-3.663.24-4.242.50-3.832.95-3.77Brachium length (BL)mean (± SD)9.29 (± 1.10)8.81 (± 1.60)8.61 (± 0.26)12.027.99 (± 0.59)8.58 (± 0.92)8.14 (± 1.04)9.12 (± 1.03)range8.51-10.066.81-10.358.32-8.98/7.08-8.856.69-9.915.78-9.567.38-10.29Forearm length (FAL)mean (± SD)10.98 (± 0.29)10.36 (± 3.36)8.07 (± 0.29)10.178.09 (± 0.40)9.70 (± 0.84)7.72 (± 0.47)8.99 (± 0.64)range10.77-11.185.79-13.947.54-8.39/7.55-8.818.87-11.397.13-8.928.23-9.92Manus length (ML)mean (± SD)13.60 (± 1.98)12.43 (± 2.86)11.03 (± 0.35)13.4110.32 (± 0.48)11.93 (± 1.04)9.67 (± 0.72)10.78 (± 0.71)range12.20-15.008.36-14.9510.37-11.53/9.65-11.1310.01-13.728.69-11.09.22-11.78Femur length (FL)mean (± SD)25.84 (± 0.37)25.29 (± 6.53)22.70 (± 0.42)27.8020.27 (± 1.26)23.83 (± 1.51)19.72 (± 1.08)21.73 (± 1.32)range25.58-26.1017.19-31.3422.2-23.22/18.38-22.0421.55-26.3717.81-21.5719.13-23.68Crus length (CL)mean (± SD)27.04 (± 3.25)26.71 (± 7.19)23.60 (± 0.42)28.7721.98 (± 0.99)25.94 (± 1.41)20.88 (± 0.70)23.20 (± 1.58)range24.74-29.3317.28-32.5022.83-41.01/20.75-23.7323.88-28.7519.50-22.0619.83-25.02Tarsal length (TL)mean (± SD)14.06 (± 1.31)12.99 (± 3.34)11.28 (± 0.40)14.4211.12 (± 0.78)13.18 (± 0.90)10.84 (± 0.50)12.35 (± 0.76)range13.13-14.988.52-16.0010.70-11.84/10.00-12.3512.25-14.569.86-11.7511.21-13.48Pes length (PL)mean (± SD)25.72 (± 3.22)24.87 (± 7.13)23.31 (± 0.79)28.8421.35 (± 1.09)25.20 (± 1.75)20.28 (± 0.88)22.62 (± 0.90)range23.44-28.0014.72-31.4922.10-24.46/19.62-23.1122.52-27.8618.40-21.8221.16-23.73Humeral gland (HG)mean (± SD)3.57 (± 0.45)/2.58 (± 0.21)/3.39 (± 0.54)/3.26 (± 0.66)/range3.25-3.89/2.40-2.98/2.58-4.41/2.18-4.16/Pattern of venterBelly usually mottled with black and white reticulations along the flanks; limbs with bold black and white bands or reticulationsSpottedImmaculate to very faint, sparse specklingImmaculate to very faint, sparse speckling表5. 棕蛙属（Abavorana）类群的汇总统计与腹面色彩模式。本表格汇总了马来半岛与婆罗洲的装饰棕蛙（Abavorana decorata）、纳兹古尔棕蛙（A. nazgul）以及哀棕蛙（A. luctuosa）种群的形态测量汇总统计数据与腹面色彩模式，其中SD为标准差（standard deviation），N为样本量（sample size），符号“/”代表无有效数据。仅雄性个体具有肱腺（humeral gland），所有形态测量指标的单位均为毫米（mm）。 |类群|性别|N|吻肛长（Snout-vent length, SVL）均值（±标准差）|吻肛长范围|头长（Head length, HL）均值（±标准差）|头长范围|头宽（Head width, HW）均值（±标准差）|头宽范围|吻长（Snout length, SL）均值（±标准差）|吻长范围|吻宽（Snout width, SW）均值（±标准差）|吻宽范围|眼间距（Inter orbital distance, IOD）均值（±标准差）|眼间距范围|鼻间距（Internarial distance, IND）均值（±标准差）|鼻间距范围|眼径（Eye diameter, ED）均值（±标准差）|眼径范围|鼓膜径（Tympanum diameter, TD）均值（±标准差）|鼓膜径范围|臂长（Brachium length, BL）均值（±标准差）|臂长范围|前臂长（Forearm length, FAL）均值（±标准差）|前臂长范围|手长（Manus length, ML）均值（±标准差）|手长范围|股骨长（Femur length, FL）均值（±标准差）|股骨长范围|胫长（Crus length, CL）均值（±标准差）|胫长范围|跗长（Tarsal length, TL）均值（±标准差）|跗长范围|足长（Pes length, PL）均值（±标准差）|足长范围|肱腺（Humeral gland, HG）均值（±标准差）|肱腺范围|腹面色彩模式| |---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---| |装饰棕蛙（A. decorata）|雄性|26|51.11（±6.92）|46.21-56.00|19.10（±0.06）|19.05-19.14|18.60（±1.32）|17.66-19.53|7.99（±0.33）|7.76-8.22|10.31（±0.57）|9.91-10.71|5.67（±0.40）|5.59-5.95|6.13（±0.05）|6.09-6.16|5.75（±0.26）|5.56-5.93|4.57（±0.04）|4.54-4.60|9.29（±1.10）|8.51-10.06|10.98（±0.29）|10.77-11.18|13.60（±1.98）|12.20-15.00|25.84（±0.37）|25.58-26.10|27.04（±3.25）|24.74-29.33|14.06（±1.31）|13.13-14.98|25.72（±3.22）|23.44-28.00|3.57（±0.45）|3.25-3.89|腹部通常沿体侧布有黑白网状斑纹；四肢具醒目的黑白环纹或网状斑纹| |装饰棕蛙（A. decorata）|雌性|7|48.73（±13.32）|29.66-58.42|18.95（±4.85）|12.23-23.71|17.18（±4.68）|10.62-21.07|7.73（±1.85）|4.69-9.46|9.54（±2.28）|6.25-11.68|5.63（±1.22）|3.91-6.78|5.76（±1.34）|3.68-6.77|5.31（±1.10）|3.94-6.58|4.30（±1.43）|2.27-5.63|8.81（±1.60）|6.81-10.35|10.36（±3.36）|5.79-13.94|12.43（±2.86）|8.36-14.95|25.29（±6.53）|17.19-31.34|26.71（±7.19）|17.28-32.50|12.99（±3.34）|8.52-16.00|24.87（±7.13）|14.72-31.49|/|/|具斑点| |纳兹古尔棕蛙（A. nazgul）|雄性|11|42.77（±1.22）|41.51-44.10|16.61（±0.25）|16.36-17.14|15.16（±0.46）|14.63-15.83|7.31（±0.20）|7.10-7.57|8.26（±0.30）|7.98-8.70|4.20（±0.23）|3.94-4.48|5.64（±0.13）|5.45-5.80|4.17（±0.24）|3.86-4.56|3.72（±0.19）|3.52-4.01|8.61（±0.26）|8.32-8.98|8.07（±0.29）|7.54-8.39|11.03（±0.35）|10.37-11.53|22.70（±0.42）|22.2-23.22|23.60（±0.42）|22.83-41.01|11.28（±0.40）|10.70-11.84|23.31（±0.79）|22.10-24.46|2.58（±0.21）|2.40-2.98|具斑点| |纳兹古尔棕蛙（A. nazgul）|雌性|1|52.51|/|19.65|/|18.57|/|8.69|/|9.59|/|4.94|/|6.61|/|4.80|/|4.10|/|12.02|/|10.17|/|13.41|/|27.80|/|28.77|/|14.42|/|28.84|/|/|/|具斑点| |马来半岛哀棕蛙（A. luctuosa from Malay Peninsula）|雄性|12|40.74（±2.54）|37.20-44.27|15.64（±0.81）|14.45-16.69|14.24（±0.66）|13.35-15.58|6.88（±0.43）|6.20-7.56|8.08（±0.44）|7.11-8.62|4.35（±0.52）|3.25-5.14|5.33（±0.30）|4.93-5.88|4.25（±0.26）|3.80-4.70|3.30（±0.31）|2.62-3.66|7.99（±0.59）|7.08-8.85|8.09（±0.40）|7.55-8.81|10.32（±0.48）|9.65-11.13|20.27（±1.26）|18.38-22.04|21.98（±0.99）|20.75-23.73|11.12（±0.78）|10.00-12.35|21.35（±1.09）|19.62-23.11|3.39（±0.54）|2.58-4.41|无斑纹，或仅具极微弱、稀疏的细点| |马来半岛哀棕蛙（A. luctuosa from Malay Peninsula）|雌性|2|47.38（±2.36）|43.89-51.06|17.72（±1.39）|15.20-19.59|15.77（±1.01）|13.83-16.84|7.82（±0.56）|6.48-8.30|8.84（±0.40）|8.21-9.38|4.93（±0.54）|3.80-5.64|6.06（±0.38）|5.42-6.72|4.87（±0.29）|4.38-5.21|3.87（±0.27）|3.24-4.24|8.58（±0.92）|6.69-9.91|9.70（±0.84）|8.87-11.39|11.93（±1.04）|10.01-13.72|23.83（±1.51）|21.55-26.37|25.94（±1.41）|23.88-28.75|13.18（±0.90）|12.25-14.56|25.20（±1.75）|22.52-27.86|/|/|无斑纹，或仅具极微弱、稀疏的细点| |婆罗洲哀棕蛙（A. luctuosa from Borneo）|雄性|11|38.15（±1.45）|35.05-40.55|15.25（±0.78）|13.53-16.78|13.39（±0.52）|12.25-14.41|6.55（±0.34）|5.66-7.03|7.73（±0.23）|7.33-8.22|4.24（±0.30）|3.78-4.70|5.16（±0.24）|4.66-5.58|4.18（±0.26）|3.65-4.59|3.16（±0.29）|2.50-3.83|8.14（±1.04）|5.78-9.56|7.72（±0.47）|7.13-8.92|9.67（±0.72）|8.69-11.09|19.72（±1.08）|17.81-21.57|20.88（±0.70）|19.50-22.06|10.84（±0.50）|9.86-11.75|20.28（±0.88）|18.40-21.82|3.26（±0.66）|2.18-4.16|无斑纹，或仅具极微弱、稀疏的细点| |婆罗洲哀棕蛙（A. luctuosa from Borneo）|雌性|1|43.23（±2.51）|37.05-45.20|16.88（±0.92）|14.84-17.72|14.69（±0.83）|12.71-15.74|7.17（±0.33）|6.61-7.86|8.33（±0.51）|7.65-9.02|4.69（±0.37）|4.15-5.60|5.50（±0.42）|4.75-5.99|4.69（±0.29）|4.08-5.22|3.44（±0.25）|2.95-3.77|9.12（±1.03）|7.38-10.29|8.99（±0.64）|8.23-9.92|10.78（±0.71）|9.22-11.78|21.73（±1.32）|19.13-23.68|23.20（±1.58）|19.83-25.02|12.35（±0.76）|11.21-13.48|22.62（±0.90）|21.16-23.73|/|/|无斑纹，或仅具极微弱、稀疏的细点|