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Table 4. A in Three new species and one subspecies of the Amynthas corticis - group from Guangxi Zhuang Autonomous Region, China (Oligochaeta, Megascolecidae)

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Table 4. A comparison of characters of A. shengtangmontis shengtangmontis, A. shengtangmontis minusculus, A. carnosus, A. corticis. and A. pulvinus. CharacteristicsA. shengtangmontis shengtangmontisA. shengtangmontis minusculusA. carnosus (Goto & Hatai, 1899)A. corticis (Kinberg, 1867)A. pulvinus Sun & Qiu, 2013Body length (mm)100-13475-83110-34045-17093.5Body width (mm)4.2-5.13.0-3.24.0-9.03.0-6.03.4PigmentdorsumFrom brown to noPurple brownDark brown or purpleGreenish brownBuffventrumFrom brown to noNoDark brown or purpleNoNoFirst dorsal pore12/1311/1212/1310/11 or 12/13, usually at 11/1212/13Clitellum localityXIV–XVIXIV–XVIXIV–XVIXIV–XVI, occasionally shorterXIV–XVISpermathecal pores4 pairs, in 5/6-8/9, 0.40C4 pairs, in 5/6-8/9, 0.40C4 pairs in 5/6-8/9 or 3 pairs in 6/7-8/9, 0.33C4 pairs, 5/6-8/9, 0.33C4 pairs, in 5/6-8/9, 0.33CMale poresLarge raised, round, surrounded by 2 circular ridges, 0.44CRaised, elliptic, surrounded by 6 circular ridges, 0.40CRound or ellipticSmall, circular to transverse elliptical disc, 0.24-0.30CSlightly elevated round, 0.33CPapillae preclitellumFour pairs of postsetal genital papillae arranged in VI–IXThree pairs of postsetal genital papillae arranged in VI–VIII8 papillae just overhead 8 spermathecal pores, 2 pairs preclitellar arranged on VIII and IXPaired presetal and/or postsetal in some or all, near spermathecal poresInvisiblePapillae postclitellum2 presetal indented-topped genital medial of porophore2 small indented-topped genital papillae medial of male pore2 paired presetal genital on XVIII and IX, 1 pair postsetal genital on XVIIIPresent or absent, occasionally one or more near male poreRectangle-shaped, on 17/18-18/19Prostate glandsXV–XXIIXVI–XXIWell developedXVII–XX, rudimentary or absentXVII–XXSpermathecaeAbout 2.2-3.0mm long, ampulla of the first three pairs heart-shaped, duct stalk. Ampulla of the forth pair long-oval, duct as long as 1/6 ampullaAbout 2.2-2.7mm long, duct as long as 1/2 ampullaAmpulla oval or pear-shaped, duct equal to or slightly shorter than ampullaAmpulla ovoidAbout 2.4mm long, ampulla slender heart-shaped; duct shortDiverticulumAs long as main chain, U-shaped twisted in middle, terminal 4/5, club-shaped seminal chamber of the first three pairs and terminal 1/3 dilated into a chilli-shaped seminal chamber of the forth pairAs long as main chain, terminal 1/2, long club-shaped seminal chamberOne-third to half of ampulla, slender stalk and a wider seminal chamberBlunt ovioid, straight stalkShorter, terminal 1/5, small ovoid plump seminal chamberAccessory glands1 round semitransparent accessory gland present near the medial area of each spermatheca, 2 stalk accessory glands observed near the medial of the distal part of the prostatic duct6 semitransparent elliptic accessory glands observed near the distal part of the last three pairs spermathecae-Stalked, coelomic, bound down to parietes or retained within body wallA pair, cling to body wall, irregular in shape, and extended from XVII–XIX

表4 圣唐山蚓指名亚种(A. shengtangmontis shengtangmontis)、圣唐山蚓小型亚种(A. shengtangmontis minusculus)、肉质蚓(A. carnosus (Goto & Hatai, 1899))、皮栖蚓(A. corticis (Kinberg, 1867))与垫状蚓(A. pulvinus Sun & Qiu, 2013)的特征比较 1. 体长(mm):圣唐山蚓指名亚种100~134,圣唐山蚓小型亚种75~83,肉质蚓110~340,皮栖蚓45~170,垫状蚓93.5 2. 体宽(mm):圣唐山蚓指名亚种4.2~5.1,圣唐山蚓小型亚种3.0~3.2,肉质蚓4.0~9.0,皮栖蚓3.0~6.0,垫状蚓3.4 3. 体色 - 背面:圣唐山蚓指名亚种从褐色至无色素,圣唐山蚓小型亚种为紫褐色,肉质蚓为深褐色或紫色,皮栖蚓为绿褐色,垫状蚓为米黄色 - 腹面:圣唐山蚓指名亚种从褐色至无色素,圣唐山蚓小型亚种无色素,肉质蚓为深褐色或紫色,皮栖蚓无色素,垫状蚓无色素 4. 首个背孔:圣唐山蚓指名亚种为12/13节段,圣唐山蚓小型亚种为11/12节段,肉质蚓为12/13节段,皮栖蚓为10/11或12/13节段(通常为11/12节段),垫状蚓为12/13节段 5. 环带位置:圣唐山蚓指名亚种为XIV~XVI节,圣唐山蚓小型亚种为XIV~XVI节,肉质蚓为XIV~XVI节,皮栖蚓为XIV~XVI节(偶有缩短),垫状蚓为XIV~XVI节 6. 受精囊孔:圣唐山蚓指名亚种4对,位于5/6~8/9节段,孔径约为体宽的0.40倍;圣唐山蚓小型亚种4对,位于5/6~8/9节段,孔径约为体宽的0.40倍;肉质蚓4对位于5/6~8/9节段或3对位于6/7~8/9节段,孔径约为体宽的0.33倍;皮栖蚓4对,位于5/6~8/9节段,孔径约为体宽的0.33倍;垫状蚓4对,位于5/6~8/9节段,孔径约为体宽的0.33倍 7. 雄孔:圣唐山蚓指名亚种为大型隆起圆形雄孔,周围环绕2圈环状脊,孔径约为体宽的0.44倍;圣唐山蚓小型亚种为隆起椭圆形雄孔,周围环绕6圈环状脊,孔径约为体宽的0.40倍;肉质蚓为圆形或椭圆形;皮栖蚓为小型圆形至横椭圆形盘状雄孔,孔径约为体宽的0.24~0.30倍;垫状蚓为轻微隆起的圆形雄孔,孔径约为体宽的0.33倍 8. 环前生殖乳突:圣唐山蚓指名亚种具4对刚毛后生殖乳突,分布于VI~IX节;圣唐山蚓小型亚种具3对刚毛后生殖乳突,分布于VI~VIII节;肉质蚓具8个乳突紧邻8个受精囊孔,另有2对环前生殖乳突位于VIII和IX节;皮栖蚓部分或全部个体具成对刚毛前/后生殖乳突,紧邻受精囊孔;垫状蚓不可见 9. 环后生殖乳突:圣唐山蚓指名亚种具2个具凹陷顶端的生殖内侧乳突,位于雄孔孔区;圣唐山蚓小型亚种具2个具凹陷顶端的小型生殖乳突,位于雄孔内侧;肉质蚓具2对刚毛前生殖乳突位于XVIII与IX节,1对刚毛后生殖乳突位于XVIII节;皮栖蚓乳突存在或缺失,偶有1个或多个乳突紧邻雄孔;垫状蚓为矩形,位于17/18~18/19节段 10. 前列腺腺:圣唐山蚓指名亚种为XV~XXII节,圣唐山蚓小型亚种为XVI~XXI节,肉质蚓发育良好,皮栖蚓为XVII~XX节(退化或缺失),垫状蚓为XVII~XX节 11. 受精囊:圣唐山蚓指名亚种长约2.2~3.0mm,前3对受精囊的壶腹呈心形,具管柄;第4对受精囊的壶腹呈长椭圆形,管长约为壶腹的1/6;圣唐山蚓小型亚种长约2.2~2.7mm,管长约为壶腹的1/2;肉质蚓的壶腹呈椭圆形或梨形,管长等于或略短于壶腹;皮栖蚓的壶腹呈卵圆形;垫状蚓长约2.4mm,壶腹呈细长心形,管短 12. 受精囊盲囊:圣唐山蚓指名亚种与主链等长,中部呈U形扭曲,前3对的末端4/5处为棒状储精室,第4对的末端1/3处膨大呈辣椒状储精室;圣唐山蚓小型亚种与主链等长,末端1/2处为长棒状储精室;肉质蚓长度为壶腹的1/3~1/2,具细长管柄和较宽的储精室;皮栖蚓为钝卵圆形,管直;垫状蚓较短,末端1/5处具小型卵圆形饱满储精室 13. 副腺:圣唐山蚓指名亚种每个受精囊内侧区域附近具1个圆形半透明副腺,前列腺管远端内侧可见2个具柄副腺;圣唐山蚓小型亚种最后3对受精囊的远端附近可见6个半透明椭圆形副腺;肉质蚓具柄,体腔来源,附着于体壁或保留于体壁内;皮栖蚓具1对,附着于体壁,形状不规则,分布于XVII~XIX节;垫状蚓具1对,附着于体壁,形状不规则,分布于XVII~XIX节

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