TABLE 1 in Description of a new species of the earthworm genus Moniligaster Perrier, 1872 (Clitellata, Moniligastridae) from the Nilgiri Biosphere Reserve of Tamil Nadu State, India

TABLE 1. Comparison of the characters of M. gatesi Ahmed & Narayanan, sp. nov., with other species of the ‘ gravelyi ’ group. CharacterM. gravelyi Stephenson, 19151,2M. troyi Jamieson, 19773M. bahli Narayanan & Julka,20214M. blakemorei Narayanan & Julka, 20214,5M. keralensis Narayanan & Julka, 20214,5M. julkai Narayanan & Paliwal, 20225M. gatesi Ahmed & Narayanan, sp. nov.Length130–174 mm60–63 mm75–134 mm81–136 mm58–155 mm156–238 mm87–160 mmDiameter5–7 mm3.5–3.7 mm4.5–7 mm3.5–7 mm3–6 mm6.5–8.5 mm5–7 mmSegments139–189123–153135–200183–278109–201237–342116–181Spermathecal pore locationbetween bc setal lines, median to setae cin c setal linebetween bc, median to setae cclose to cd setal linesaligned between cd setal linesaligned at cd setal linesbetween bc setal lines, nearer to cClitellumannular, on segments ¼9– ½14 (= ¾4)annular, on segments ½9–14 (=5½)annular, on segments 10–13 (=4)annular, on segments 10–13 (=4)annular, on segments 10–13 (=4)annular, on segments 10–½14 (=4½)annular, on segments ⅓9–14 (=5⅓)Secondary male aperture or male pore locationaligned between bc setal lines, nearer to bon evaginated long cylindrical pseudopenes, at bc setal lines, lateral to blateral to seta b linesslightly lateral to b setal lineslightly lateral to b setal lineslateral to b setal linebetween bc setal lines, nearer to bFemale pore locationat b setal line?minute, at b setal lineminute, at b setal lineminute, slightly median to b setal lineminute, at b setal line?b setal lineSpermathecaatrial gland single, on each side; duct of the gland c. 1/7 the length of common atrial duct; ampulla ovoidatrial glands paired on each side; duct of each gland c. 1/4 the length of common atrial duct; ampulla elongate-ovoidatrial glands paired on each side; duct of each gland c. 1/5 the length of common atrial duct; ampulla ovoidatrial glands paired on each side; duct of each gland c. four times the length of common atrial duct; ampulla ovoidalatrial glands paired on each side; duct of each gland about as long as common atrial duct; ampulla elongate-ovoidatrial glands paired on each side; duct of each gland c. five times the length of common atrial duct; ampulla ovoidalatrial glands paired on each side; duct of the gland c. 1/6 the length of common atrial duct; ampulla pear-shapedProstatesstrap-shapedclub-shapedstrap-shapedbluntly club-shapedtubulartubularstrap-shapedProstate capsuletubular with deeply incised margins and highly nodulated surface(?)tubular with smooth margins and a few nodulations at ental endclub-shaped with smooth marginstubular with smooth marginstubular with smooth marginstubular, surface with shallow nodulations, margins nodulatedLength of prostatic ductjust over 1/4 the length of gland plus ductc.1/3 the length of gland plus ductc. 1/7 the length of gland plus ductc. 1/3 the length of gland plus ductc. 1/9 the length of gland plus ductc. 1/3 the length of gland plus ductc. 1/8 the length of gland plus ductVas deferenssegments 9–10, 11, enters prostate sub-entallysegment 9, enters prostate sub-ectallysegments 9–12(–16), enters prostate sub-entallysegments 9–11(– 12), enters prostate near entallysegments 9–10, enters prostate sub-ectallysegments 9–12(–13), enters prostate sub-ectallysegments 9–10, enters prostate sub-entallyGizzards4–5 (in segments 12–18)3 (in segments 13–15)3–4 (in segments 13–18)2 (in segments 12–16)3–6(in segments 14–20)2–3(in segments 12–16)4–5 (in segments 13–18)Intestinal originin segment 25±1in segment 27in segment 25±1in segment 27±1in segment 28±1in segment 27in segment 26±1Ovisacsextending to segments 13–14extending to segment 13extending to segment 20extending to segment 13extending to segments 12–16extending to segments 13–14extending to segments 13–16 ......continued on the next page Data from: 1 Stephenson (1915); 2 Narayanan et al. (2023b); 3 Jamieson (1977); 4 Narayanan et al. (2021a); 5 Narayanan et al. (2022).表1 盖茨氏新种（*M. gatesi* Ahmed & Narayanan, sp. nov.）与"gravelyi"类群其他物种的形态特征对比 | 形态特征 | *M. gravelyi* Stephenson, 1915¹,² | *M. troyi* Jamieson, 1977³ | *M. bahli* Narayanan & Julka, 2021⁴ | *M. blakemorei* Narayanan & Julka, 2021⁴,⁵ | *M. keralensis* Narayanan & Julka, 2021⁴,⁵ | *M. julkai* Narayanan & Paliwal, 2022⁵ | *M. gatesi* Ahmed & Narayanan, sp. nov. | | --- | --- | --- | --- | --- | --- | --- | --- | | 体长 | 130–174 mm | 60–63 mm | 75–134 mm | 81–136 mm | 58–155 mm | 156–238 mm | 87–160 mm | | 体宽 | 5–7 mm | 3.5–3.7 mm | 4.5–7 mm | 3.5–7 mm | 3–6 mm | 6.5–8.5 mm | 5–7 mm | | 体节数 | 139–189 | 123–153 | 135–200 | 183–278 | 109–201 | 237–342 | 116–181 | | 受精囊孔位置 | bc刚毛列之间，c刚毛列内侧 | 位于c刚毛列 | bc刚毛列之间，c刚毛列内侧 | 靠近cd刚毛列 | 对齐于cd刚毛列之间 | 对齐于cd刚毛列上 | bc刚毛列之间，更靠近c | | 环带 | 环状，位于第9¼至14½体节（即4¾体节段） | 环状，位于第9½至14体节（即5½体节段） | 环状，位于第10–13体节（即4体节段） | 环状，位于第10–13体节（即4体节段） | 环状，位于第10–13体节（即4体节段） | 环状，位于第10至14½体节（即4½体节段） | 环状，位于第9⅓至14体节（即5⅓体节段） | | 次生雄孔/雄孔位置 | 对齐于bc刚毛列之间，更靠近b | 着生于外翻的长圆柱形假阴茎上，位于bc刚毛列，靠近b外侧 | 位于b刚毛列外侧 | 略位于b刚毛列外侧 | 略位于b刚毛列外侧 | 位于b刚毛列外侧 | 对齐于bc刚毛列之间，更靠近b | | 雌孔位置 | 位于b刚毛列 | 微小，位于b刚毛列 | 微小，位于b刚毛列 | 微小，略偏向b刚毛列内侧 | 微小，位于b刚毛列 | 未知 | 位于b刚毛列 | | 受精囊 | 单侧具单个心房腺；腺导管长约为总心房导管的1/7；受精囊坛呈卵形 | 单侧具成对心房腺；各腺导管长约为总心房导管的1/4；受精囊坛呈长卵形 | 单侧具成对心房腺；各腺导管长约为总心房导管的1/5；受精囊坛呈卵形 | 单侧具成对心房腺；各腺导管长约为总心房导管的4倍；受精囊坛呈卵形 | 单侧具成对心房腺；各腺导管长约与总心房导管相当；受精囊坛呈长卵形 | 单侧具成对心房腺；各腺导管长约为总心房导管的5倍；受精囊坛呈卵形 | 单侧具成对心房腺；腺导管长约为总心房导管的1/6；受精囊坛呈梨形 | | 前列腺 | 带状 | 棒状 | 带状 | 钝棒状 | 管状 | 管状 | 带状 | | 前列腺囊 | 管状，边缘深裂，表面高度结节化 | ？管状，边缘光滑，近端具少量结节 | 棒状，边缘光滑 | 管状，边缘光滑 | 管状，边缘光滑 | 管状，表面具浅结节，边缘结节化 | 未明确标注 | | 前列腺导管长度 | 略长于腺体加导管总长的1/4 | 约为腺体加导管总长的1/3 | 约为腺体加导管总长的1/7 | 约为腺体加导管总长的1/3 | 约为腺体加导管总长的1/9 | 约为腺体加导管总长的1/3 | 约为腺体加导管总长的1/8 | | 输精管 | 跨越第9–10、11体节，于前列腺近端通入 | 跨越第9体节，于前列腺远端通入 | 跨越第9–12（–16）体节，于前列腺近端通入 | 跨越第9–11（–12）体节，近前列腺近端通入 | 跨越第9–10体节，于前列腺远端通入 | 跨越第9–12（–13）体节，于前列腺远端通入 | 跨越第9–10体节，于前列腺近端通入 | | 砂囊 | 4–5个，位于第12–18体节 | 3个，位于第13–15体节 | 3–4个，位于第13–18体节 | 2个，位于第12–16体节 | 3–6个，位于第14–20体节 | 2–3个，位于第12–16体节 | 4–5个，位于第13–18体节 | | 肠起始节 | 第25±1体节 | 第27体节 | 第25±1体节 | 第27±1体节 | 第28±1体节 | 第27体节 | 第26±1体节 | | 卵囊 | 延伸至第13–14体节 | 延伸至第13体节 | 延伸至第20体节 | 延伸至第13体节 | 延伸至第12–16体节 | 延伸至第13–14体节 | 延伸至第13–16体节 | ……续下页 数据来源：1 Stephenson（1915）；2 Narayanan等（2023b）；3 Jamieson（1977）；4 Narayanan等（2021a）；5 Narayanan等（2022）。