Table 5 in Taxonomic Status of Nanotyrannus lancensis (Dinosauria: Tyrannosauroidea) - A Distinct Taxon of Small-Bodied Tyrannosaur

Table 5. LAG data, body mass, and growth rate estimates rounded to the nearest kg for Sue (FMNH PR 2081), Petey (BMRP 2006.4.4), and Jane (BMRP 2002.4.4) based on femoral circumferences, including corrected values for Jane based on a split multi-LAG interpretation [49]. Campione et al. [50] estimate Sue’s mass at death (i.e., after LAG #23) as 7377 kg after applying a correction for non-circular femoral cross-section. Hutchinson et al. [104] estimate Jane’s mass at death (i.e., after LAG #9) as 954 kg based on 3D modeling. See Supplemental File for equations used in calculations. Note that LAG #1 and the periosteum do not record a full year of growth. Periosteum is assigned a LAG pseudo-number of +0.5 after the final LAG, indicating that death could have occurred at any point after the last yearly marker was deposited while assigning an age at the midpoint of that final year. Jane required estimation of LAG/periosteum circumferences from LAG spacing and femoral width due to the incomplete cross-section of the femur midshaft. (1) LAG (or periosteum) distance from endosteum is measured at the time of death, (2) femoral width at midshaft (including medullary cavity) is measured at time of death and back estimated using LAG spacing, (3) femoral circumference derived from femoral width at a given year is approximated as a circular cross-section. Data sources: Campione et al. [50]; Cullen et al. [49]; Woodward et al. [46]. SpecimenNicknameTaxonLAG No.Circumference (mm)Mass (kg) (Campione et al., 2014), Equation (7))Mass Increase from Previous LAG (kg)FMNH PR 2081SueTyrannosaurus1338.71924NAFMNH PR 2081SueTyrannosaurus23872777853FMNH PR 2081SueTyrannosaurus3415.73382605FMNH PR 2081SueTyrannosaurus4440.33962580FMNH PR 2081SueTyrannosaurus5459.74461500FMNH PR 2081SueTyrannosaurus6485.55185724FMNH PR 2081SueTyrannosaurus7511.15973788FMNH PR 2081SueTyrannosaurus8527.16503529FMNH PR 2081SueTyrannosaurus9542.87050547FMNH PR 2081SueTyrannosaurus10551.67369319FMNH PR 2081SueTyrannosaurus11559.37656287FMNH PR 2081SueTyrannosaurus12564.97869213FMNH PR 2081SueTyrannosaurus13565.7790031FMNH PR 2081SueTyrannosaurus14566.5793131FMNH PR 2081SueTyrannosaurus15567.4796535FMNH PR 2081SueTyrannosaurus16568.2799631FMNH PR 2081SueTyrannosaurus17569802731FMNH PR 2081SueTyrannosaurus18569.8805931FMNH PR 2081SueTyrannosaurus19570.7809435FMNH PR 2081SueTyrannosaurus20571.5812531FMNH PR 2081SueTyrannosaurus21572.3815631FMNH PR 2081SueTyrannosaurus22573.1818831FMNH PR 2081SueTyrannosaurus23574822335FMNH PR 2081SueTyrannosaurusPeriosteum5798422199BMRP 2006.4.4PeteyNanotyrannus1148.4198NABMRP 2006.4.4PeteyNanotyrannus2165.926971BMRP 2006.4.4PeteyNanotyrannus3194.7419149BMRP 2006.4.4PeteyNanotyrannus4214.5547128BMRP 2006.4.4PeteyNanotyrannus5223.661366BMRP 2006.4.4PeteyNanotyrannus6229.465845BMRP 2006.4.4PeteyNanotyrannus7234.870144BMRP 2006.4.4PeteyNanotyrannusPeriosteum244.778684BMRP 2002.4.4JaneNanotyrannus1195.9426NABMRP 2002.4.4JaneNanotyrannus2218.4575149BMRP 2002.4.4JaneNanotyrannus3231.8677102BMRP 2002.4.4JaneNanotyrannus4247.3809132BMRP 2002.4.4JaneNanotyrannus5248.682011BMRP 2002.4.4JaneNanotyrannus6255.388463BMRP 2002.4.4JaneNanotyrannus7257.790623BMRP 2002.4.4JaneNanotyrannus8265.798679BMRP 2002.4.4JaneNanotyrannus9266.69959BMRP 2002.4.4JaneNanotyrannusPeriosteum273.2106469BMRP 2002.4.4Jane (corrected)Nanotyrannus1195.9426NABMRP 2002.4.4Jane (corrected)Nanotyrannus2218.4575149BMRP 2002.4.4Jane (corrected)Nanotyrannus3231.8677102BMRP 2002.4.4Jane (corrected)Nanotyrannus4248.6820144BMRP 2002.4.4Jane (corrected)Nanotyrannus5257.790686BMRP 2002.4.4Jane (corrected)Nanotyrannus6266.699588BMRP 2002.4.4Jane (corrected)NanotyrannusPeriosteum273.2106469表5。基于股骨周长的生长停滞线（LAG）数据、体重及生长速率估算值，所有数值均取整至最接近的千克数，涉及标本Sue（FMNH PR 2081）、Petey（BMRP 2006.4.4）与Jane（BMRP 2002.4.4），其中Jane的估算值基于拆分式多生长停滞线解释方法得到的校正数据[49]。Campione等人[50]在对非圆形股骨横截面进行校正后，估算出Sue死亡时（即第23条生长停滞线之后）的体重为7377千克。Hutchinson等人[104]基于三维建模方法，估算出Jane死亡时（即第9条生长停滞线之后）的体重为954千克。计算所用的公式详见补充文件。需注意，第1条生长停滞线与骨膜并未记录完整一年的生长情况。研究人员将最终生长停滞线之后的骨膜标记为伪生长停滞线编号+0.5，这意味着死亡可能发生在最后一条年度生长标记形成后的任意时刻，同时将死亡年龄设定为该最后年度生长周期的中点。由于Jane的股骨中轴横截面不完整，需通过生长停滞线间距与股骨宽度估算其生长停滞线/骨膜周长。具体步骤如下：(1) 测量死亡时刻的生长停滞线（或骨膜）与骨内膜之间的距离；(2) 测量死亡时刻的股骨中轴宽度（包含髓腔），并通过生长停滞线间距反向估算各时期的股骨宽度；(3) 将给定年份的股骨宽度近似为圆形横截面，由此推导股骨周长。数据来源：Campione等人[50]；Cullen等人[49]；Woodward等人[46]。 表5详细数据如下： | 标本编号 | 昵称 | 分类群 | 生长停滞线编号 | 周长（mm） | 体重（kg，Campione等人2014年研究，公式(7)） | 相较上一条生长停滞线的体重增量（kg） | |---------|------|--------|----------------|------------|--------------------------------------------|----------------------------------------| | FMNH PR 2081 | Sue | 霸王龙（Tyrannosaurus） | 1 | 338.7 | 1924 | NA | | FMNH PR 2081 | Sue | 霸王龙 | 2 | 387 | 2777 | 853 | | FMNH PR 2081 | Sue | 霸王龙 | 3 | 415.7 | 3382 | 605 | | FMNH PR 2081 | Sue | 霸王龙 | 4 | 440.3 | 3962 | 580 | | FMNH PR 2081 | Sue | 霸王龙 | 5 | 459.7 | 4461 | 500 | | FMNH PR 2081 | Sue | 霸王龙 | 6 | 485.5 | 5185 | 724 | | FMNH PR 2081 | Sue | 霸王龙 | 7 | 511.1 | 5973 | 788 | | FMNH PR 2081 | Sue | 霸王龙 | 8 | 527.1 | 6503 | 529 | | FMNH PR 2081 | Sue | 霸王龙 | 9 | 542.8 | 7050 | 547 | | FMNH PR 2081 | Sue | 霸王龙 | 10 | 551.6 | 7369 | 319 | | FMNH PR 2081 | Sue | 霸王龙 | 11 | 559.3 | 7656 | 287 | | FMNH PR 2081 | Sue | 霸王龙 | 12 | 564.9 | 7869 | 213 | | FMNH PR 2081 | Sue | 霸王龙 | 13 | 565.7 | 7900 | 31 | | FMNH PR 2081 | Sue | 霸王龙 | 14 | 566.5 | 7931 | 31 | | FMNH PR 2081 | Sue | 霸王龙 | 15 | 567.4 | 7965 | 35 | | FMNH PR 2081 | Sue | 霸王龙 | 16 | 568.2 | 7996 | 31 | | FMNH PR 2081 | Sue | 霸王龙 | 17 | 569 | 8027 | 31 | | FMNH PR 2081 | Sue | 霸王龙 | 18 | 569.8 | 8059 | 31 | | FMNH PR 2081 | Sue | 霸王龙 | 19 | 570.7 | 8094 | 35 | | FMNH PR 2081 | Sue | 霸王龙 | 20 | 571.5 | 8125 | 31 | | FMNH PR 2081 | Sue | 霸王龙 | 21 | 572.3 | 8156 | 31 | | FMNH PR 2081 | Sue | 霸王龙 | 22 | 573.1 | 8188 | 31 | | FMNH PR 2081 | Sue | 霸王龙 | 23 | 574 | 8223 | 35 | | FMNH PR 2081 | Sue | 霸王龙 | 骨膜 | 579 | 8422 | 199 | | BMRP 2006.4.4 | Petey | 矮暴龙（Nanotyrannus） | 1 | 148.4 | 198 | NA | | BMRP 2006.4.4 | Petey | 矮暴龙 | 2 | 165.9 | 2697 | | | BMRP 2006.4.4 | Petey | 矮暴龙 | 3 | 194.7 | 4191 | 1494 | | BMRP 2006.4.4 | Petey | 矮暴龙 | 4 | 214.5 | 5471 | 1280 | | BMRP 2006.4.4 | Petey | 矮暴龙 | 5 | 223.6 | 6136 | 665 | | BMRP 2006.4.4 | Petey | 矮暴龙 | 6 | 229.4 | 6584 | 448 | | BMRP 2006.4.4 | Petey | 矮暴龙 | 7 | 234.8 | 7014 | 430 | | BMRP 2006.4.4 | Petey | 矮暴龙 | 骨膜 | 244.7 | 7868 | 854 | | BMRP 2002.4.4 | Jane | 矮暴龙 | 1 | 195.9 | 426 | NA | | BMRP 2002.4.4 | Jane | 矮暴龙 | 2 | 218.4 | 575 | 149 | | BMRP 2002.4.4 | Jane | 矮暴龙 | 3 | 231.8 | 677 | 102 | | BMRP 2002.4.4 | Jane | 矮暴龙 | 4 | 247.3 | 809 | 132 | | BMRP 2002.4.4 | Jane | 矮暴龙 | 5 | 248.6 | 820 | 11 | | BMRP 2002.4.4 | Jane | 矮暴龙 | 6 | 255.3 | 884 | 64 | | BMRP 2002.4.4 | Jane | 矮暴龙 | 7 | 257.7 | 906 | 22 | | BMRP 2002.4.4 | Jane | 矮暴龙 | 8 | 265.7 | 986 | 79 | | BMRP 2002.4.4 | Jane | 矮暴龙 | 9 | 266.6 | 995 | 9 | | BMRP 2002.4.4 | Jane | 矮暴龙 | 骨膜 | 273.2 | 1064 | 69 | | BMRP 2002.4.4 | Jane（校正后） | 矮暴龙 | 1 | 195.9 | 426 | NA | | BMRP 2002.4.4 | Jane（校正后） | 矮暴龙 | 2 | 218.4 | 575 | 149 | | BMRP 2002.4.4 | Jane（校正后） | 矮暴龙 | 3 | 231.8 | 677 | 102 | | BMRP 2002.4.4 | Jane（校正后） | 矮暴龙 | 4 | 248.6 | 820 | 144 | | BMRP 2002.4.4 | Jane（校正后） | 矮暴龙 | 5 | 257.7 | 906 | 86 | | BMRP 2002.4.4 | Jane（校正后） | 矮暴龙 | 6 | 266.6 | 995 | 88 | | BMRP 2002.4.4 | Jane（校正后） | 矮暴龙 | 骨膜 | 273.2 | 1064 | 69 |