TABLE 1. Comparison of Paraputo blackmani sp. n. with other species of Paraputo with fewer than 17 distinct pairs of cerarii. in -- A- - new- - species- - of- - Paraputo- - Laing- - 1929 -- (Hemiptera: -- Coccomorpha: -- Pseudococcidae) -- from- - India
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TABLE1. Comparison of Paraputo blackmani sp.n. with other species of Paraputo with fewer than 17 distinct pairs of cerarii. SpeciesNo.pairs ofcerariiNo. conical setaein anallobe cerariusNo. antennal segsMultilocular discporedistributionTubularducts(no. typesanddistribution)Additional differences from P. blackmaniReferenceP. blackmani sp.n.727 (occas. 6)Single / double rows across posterior edges of abd. segs VI & VII, and on anterior edge of seg. VIII2 types, slightly different lengths: longer type on submargins of abd. segs VI, VII & VIII; mixed types across middles of segs V and VI; and shorter type posterior to vulva and between anal lobesn/an/aP. anomalus11–13156Confined to the mid-region of body. A group of ca. 25 posterior to vulva, and ca. 50 in transverse row on each of abd. segs VI and VII2 types; larger type in marginal groups (20‒30) on abd. segs VII & VIII; 1 or 2 in middle of seg. VII. Smaller type in rows across middles of segs VI & VII, around groups of larger type, and on anal lobe marginsAnal ring situated nearly 3x its length from apex of abdomenWilliams (1958)P. asparodosensis4157Around vulva and on abd. seg VII2 types; larger type on head and dorsal abdominal segs; smaller type on venter of posterior abdominal segsCoxa of metathoracic leg very wide, with numerous translucent poresFoldi & Kozár (2006)P. barbatus530–407–82 (occas. 3) small groups on ventral mid-region of posterior abdominal segs1 type; very few on venter of anal lobes and abd. seg. VII marginsLegs without translucent poresDe Lotto (1964)P. formicarii4–6 (and 0–2 on head)4–98Absent2 types; smaller type dorsal, normally few on abd. segs VIII & IX; sometimes a few near margin of prothorax. Larger type ventral, scattered, most abundant on posterior abd. segs, fewer anteriorlyTranslucent pores absent from hind coxae but present on tibiaeMiller & McKenzie (1973)P. gallicolus3–428Around vulva and mid-region of abdominal seg. VII1 type, in ventral mid-region of abd. segs VI & VII, submargins of seg. VIII and in a group of 2–5 between antennal basesHind coxa, femur and tibia all with numerous translucent pores. Anal lobe bar short, slender & faintMamet (1954)P. glandulifer58–96In posterior rows across mid-regions of abd. segs IV‒VII, & a group of ca. 12 posterior to vulva1 type, sparse; abd. segs V, VI & VII with 1 on each margin, and a marginal group of 2 or 3 on seg. VIIIAnal ring situated about 1x its length from posterior end of bodyWilliams (1958)P. guadalcanalae996AbsentAll narrower than a trilocular pore; medially near posterior edges of abd. segs V & VI; small marginal groups between segs VI‒VII, and VII‒VIIIAnal ring situated about 3x its length from apex of abdomenWilliams (2005)P. loranthi9178Around vulva and on 3 prevulvar segments1 type, in marginal areas of last 4 abd. segsAnal ring situated at least 2x its length from apex of abdomenStrickland (1947)P. mangiferae14Many-Next to vulvaPerhaps 1 type, large oral collar ducts on head and next to some cerariiAnal ring with multiple setae (no. variable)Williams (2004)P. markhamiae28 (occas.7)In 4 or 5 transverse groups on ventral mid- abdomen2 sizes; larger type in small groups on ventral margins of last 4 or 5 abd. segs. Smaller type very few, on abdominal venterVenter of anal lobe with elongate sclerotized anal lobe barDe Lotto (1964)P. mazoensis6–738On abd. segs IV‒IX; segs V & VI with single transverse rows; segs VII & VIII forming double transverse rows not extending to margins1 type, in transverse rows and lateral groups on abd. segs V, VI & VII; on seg. V, forming small lateral groups only, and a few present around anal lobesVenter of anal lobe with faintly sclerotized anal lobe bar and a slender bar setaWilliams (1958)P. myrmecophilus47–86In vulvar region only2 types, mixed on posterior abd. segs, amongst multilocular disc-pores in mid-region and on margins and submarginsDorsal setae small and curved, spine-likeMamet (1951)P. phoradendri6–711–197 or 8On abd. segs VIII & IX2 intergrading sizes; in clusters near cerarii on abd. segs V or VI‒IX; also medially on segs VIII, VII & sometimes VI, and in cluster near each frontal cerariusAnal ring ca. 1x its diameter from body apex, surrounded by many setae. Dorsal discoidal pores of 2 intergrading sizes, scattered; large pores in medial clusters on abd. segs VII & VIIIMiller & McKenzie (1973)P. szemaoensis5–7Many6–7Forming rows and bands across 5 posterior abdominal segs1 type, around vulva amongst multilocular disc-poresAnal ring at least 2x its length from abdominal apex; ring bearing multiple setaeDanzig & GavrilovZimin (2015)P. taquaraeContinuous band on posterior abd. segs onlyMany7Abdomen only, many, in median rows, around vulva and on seg. VI. Trilocular pores abundant1 type, across abd. segs VI & VII, few; also around margins of posterior abd. segsCerarii in almost continuous band of numerous conical setaeWilliams & Granara de WillinkP. villosus9–141–88Around vulva and mid-region of abd. segs as far forward as seg. V, 0‒2 on seg. IV1 type, in groups on submargins of abd. segs V‒VIIAnal lobe venter with irregular sclerotized patch but no anal lobe barDe Lotto (1969) ......continued on the next page ......continued on the next page
表1 副粉蚧属（Paraputo）蜡腺群（cerarii）对数少于17的物种与新种Paraputo blackmani sp. n.的比较 | 物种 | 蜡腺群对数 | 肛叶（anal lobe）蜡腺群锥形刚毛数 | 触角节（antennal segments）数 | 多室盘孔（multilocular discpore）分布 | 管状腺（tubular ducts）（类型数量与分布） | 与P. blackmani sp. n.的额外差异 | 参考文献 | | :--- | :--- | :--- | :--- | :--- | :--- | :--- | :--- | | *Paraputo blackmani* sp. n. | 7 | 27（偶见6） | - | 单/双列分布于腹节（abdominal segments，简称腹节）VI、VII后缘及腹节VIII前缘 | 2种类型，长度略有差异：长型分布于腹节VI、VII、VIII亚缘区；混合型分布于腹节V、VI中部；短型分布于阴门（vulva）后方及肛叶之间 | 无相关记录 | - | | *Paraputo anomalus* | 11–13 | 15 | 6 | 局限于体中区。阴门后方约25根，腹节VI、VII各有一横列约50根 | 2种类型：大型类型分布于腹节VII、VIII的缘群（20–30个）；腹节VII中部具1–2个。小型类型分布于腹节VI、VII中部横列、大型类型群周围及肛叶缘 | 肛环（anal ring）距腹部顶端约为其长度的3倍 | Williams (1958) | | *Paraputo asparodosensis* | 4 | 15 | 7 | 分布于阴门及腹节VII | 2种类型：大型类型分布于头部及背侧腹节；小型类型分布于后部腹节的腹面 | 后胸足基节极宽，具大量透明孔（translucent pores） | Foldi & Kozár (2006) | | *Paraputo barbatus* | 5 | 30–40 | 7–8（偶见3） | 后部腹节腹面中区具2个小群 | 1种类型，肛叶腹面及腹节VII缘分布极少 | 足无透明孔 | De Lotto (1964) | | *Paraputo formicarii* | 4–6（头部具0–2个） | 4–9 | 8 | 无 | 2种类型：小型类型为背侧型，腹节VIII、IX分布极少，有时前胸叶缘附近具少量；大型类型为腹侧型，散生，后部腹节分布最多，前部较少 | 后足基节无透明孔，但胫节具透明孔 | Miller & McKenzie (1973) | | *Paraputo gallicolus* | 3–4 | 28 | - | 分布于阴门及腹节VII中区 | 1种类型，分布于腹节VI、VII腹面中区、腹节VIII亚缘区以及触角基部之间的2–5个群 | 后足基节、股节及胫节均具大量透明孔。肛叶杆（anal lobe bar）短、纤细且模糊 | Mamet (1954) | | *Paraputo glandulifer* | 5 | 8–9 | 6 | 分布于腹节IV–VII中区的后部横列，阴门后方约12个 | 1种类型，分布稀疏；腹节V、VI、VII每缘具1个，腹节VIII具1个缘群（2–3个） | 肛环距体后端约为其长度的1倍 | Williams (1958) | | *Paraputo guadalcanalae* | 9 | 9 | 6 | 无 | 所有类型均窄于三孔盘孔（trilocular pore）；分布于腹节V、VI后缘近中区处；腹节VI–VII、VII–VIII之间具小型缘群 | 肛环距腹部顶端约为其长度的3倍 | Williams (2005) | | *Paraputo loranthi* | 9 | 17 | 8 | 分布于阴门及前3个阴前腹节 | 1种类型，分布于最后4个腹节的缘区 | 肛环距腹部顶端至少为其长度的2倍 | Strickland (1947) | | *Paraputo mangiferae* | 14 | 极多 | - | 紧邻阴门 | 约1种类型，头部及部分蜡腺群旁具大型口领腺 | 肛环具多根刚毛（数量可变） | Williams (2004) | | *Paraputo markhamiae* | 2 | 8（偶见7） | - | 腹腹中部分布有4–5个横群 | 2种尺寸：大型类型分布于最后4–5个腹节的腹侧缘小群；小型类型极少，分布于腹部腹面 | 肛叶腹面具延长的骨化（sclerotized）肛叶杆 | De Lotto (1964) | | *Paraputo mazoensis* | 6–7 | 38 | - | 分布于腹节IV–IX；腹节V、VI具单横列；腹节VII、VIII为双横列，不达缘区 | 1种类型，分布于腹节V、VI、VII的横列及侧群；腹节V仅具小型侧群，肛叶周围具少量 | 肛叶腹面具微弱骨化的肛叶杆及纤细的杆状刚毛 | Williams (1958) | | *Paraputo myrmecophilus* | 4 | 7–8 | 6 | 仅分布于阴门区 | 2种类型，混合分布于后部腹节、中区多室盘孔之间以及缘区和亚缘区 | 背侧刚毛细小且弯曲，呈刺状 | Mamet (1951) | | *Paraputo phoradendri* | 6–7 | 11–19 | 7或8 | 分布于腹节VIII、IX | 2种过渡型尺寸；簇生于腹节V或VI–IX的蜡腺群附近；亦分布于腹节VIII、VII的中区，有时腹节VI，以及每个额部蜡腺群附近 | 肛环距体顶端约为其直径的1倍，周围具多根刚毛。背侧盘状孔具2种过渡型尺寸，散生；大型孔簇生于腹节VII、VIII中区 | Miller & McKenzie (1973) | | *Paraputo szemaoensis* | 5–7 | 极多 | 6–7 | 分布于5个后部腹节的横列及带区 | 1种类型，分布于阴门周围多室盘孔之间 | 肛环距腹部顶端至少为其长度的2倍；肛环具多根刚毛 | Danzig & Gavrilov-Zimin (2015) | | *Paraputo taquarae* | 仅后部腹节具连续带 | 极多 | 7 | 仅腹部具大量孔，呈中列、阴门周围及腹节VI分布。三孔盘孔丰富 | 1种类型，分布于腹节VI、VII，数量极少；亦分布于后部腹节的缘区 | 蜡腺群呈近乎连续的多锥形刚毛带 | Williams & Granara de Willink | | *Paraputo villosus* | 9–14 | 1–8 | 8 | 分布于阴门及腹节中区，向前可达腹节V，腹节IV具0–2个 | 1种类型，簇生于腹节V–VII的亚缘区 | 肛叶腹面具不规则骨化斑，但无肛叶杆 | De Lotto (1969) | ……续下页
