Table 3. Amutatiola macroventosa n. gen. & n in Two new genera and species of sepioline squids (Cephalopoda: Sepiolidae) from Australia

Table 3. Amutatiola macroventosa n.gen. & n.sp. measurement, counts and indices of male specimens Species MuseumMOVMOVAmutatiola macroventosa WAM MOVMOVMOVMOVRegistration number F80081F800833091-83F158244F275295F80087F91362Specimen number1234567Type status CruiseHolotype SO1/85Paratype 3 SO1/85Paratype 2 Paratype 5 N. Sinclair, 4 Apr1983 –Paratype 7 –– –Paratype 8 –Station number1241044 Apr 1983, Rotenone – Station.–––SexmalemalemalemalemalemalemaleMaturity5555454DML (mm)9.89.69.58.88.186.9VMLI109.293.8102.1–88116.3103.6HLI43.947.94049.359.663.854.1HWI NCI MWI88.8 41.1 96.970.8 38.6 91.790.5 45.8 87.484.2 – –87.7 43.7 91.6101.3 40.2 115102 50.5 111.4FuLI69.462.570.5–81.261.385.5FuWI5043.842.134.655.131.358.8FFuI FLI60.3 62.266.7 58.368.7 53.7– –77.8 5461.2 63.879 64.9FBLI35.730.231.630.329.841.352.9FWI46.938.533.759.748.158.858.4A1LI A2LI A3LI92.9 87.8 73.587.5 82.3 83.3– 82.1 71.6140.9 90.9 –75.3 91 –80 98.8 88.892.4 103.6 105.2A4LI74.565.673.768.275.277.590.9Arm formula1.2.4.3.1.3.2.4.2.1.4.3.–2.1.3.4.2.3.1.4.3.2.1.4.AR1LI TtLI84.7 –91.7 161.876.8 121.1102.3 –72.8 –100 87.588.4 –ClLI24.521.929.522.732.128.826.1ClRC66666–86–86–8GLI GWI42.9 22.442.7 16.7– –– –40 25.333.7 16.346.4 18.8GiLC2121––171616A1SC21201920142816A2SC A3SC A4SC30 – 2930 – 2930 27 2732 – 2133 24 2427 – 2628 24 22AR1SC191817–122514ClSI2.41.61.90.61.62.31.9A1SI A2SI21.4 4.717.6 9.719.8 5.118.6 7.320.4 17.411.9 –24.2 6.7A3SI13.78.414.517.220.3–20.4A4SI SpLI4.1 4.8 a:36.2; b:33.5; c:35.7; d:34.0 38.88.1 42.75.6 29.36.3 –– 44.87.4 –SpWI SpRIa:6.2; b:6.7; c:5.7; d:6 a:33.5; b:35.7; c:34; d:43.26.7 40.35.4 35.56.6 37.6– –7.0 27.9– –CBIa:20.3; b:22; c:19.1; d:2019.418.519.4–10.9– Note: Maturity 4, subadult – sexual characters well distinguished but gonads and accessory organs not completely developed; Maturity 5, adult – sexually mature with spermatophores in Needham’s sac in males; ovaries, nidamental and oviducal glands fully developed and ripe, eggs sometimes in oviducts, in females; DML, dorsal mantle length (mm) – measured from anterior-most point of mantle to posterior end of mantle; VMLI, ventral mantle length index – ventral mantle length, measured from anterior border of mantle at ventral midline, to apex of mantle, expressed as a percentage of DML; HLI, head length index – dorsal length of head measured from point of fusion of dorsal arms to anterior tip of dorsal mantle expressed as a percentage of DML; HWI, head width index – greatest width of head at level of eyes expressed as a percentage of DML; NCI, nuchal commissure width index – width of nuchal commissure expressed as a percentage of width of mantle at the position of nuchal commissure; MWI, mantle width index – mantle width at mantle opening expressed as a percentage of DML; FuLI, funnel length index – length of funnel from anterior funnel opening to posterior border of funnel measured along ventral midline expressed as a percentage of DML; FuWI, funnel width index – width of funnel at junction of funnel and head just anterior to funnel locking cartilages expressed as a percentage of DML; FFuI, free funnel index – length of funnel from the anterior opening to the point of dorsal attachment to head expressed as a percentage of funnel length; FLI, fin length index – greatest length of a single fin expressed as a percentage of DML; FBLI, fin base length index – length of fin attachment to mantle expressed as a percentage of DML; FWI, fin width index – greatest width of a single fin expressed as a percentage of DML; A 1 LI, arm I length index – length of arm I expressed as a percentage of DML; A 2 LI, arm II length index – length of arm II expressed as a percentage of DML; A LI, arm III length index – length of arm III expressed as a percentage of DML; A LI, arm IV length index – length of arm IV 3 4 expressed as a percentage of DML; A LI, right arm I length index – length of right arm I expressed as a percentage of DML (only in males); TtLI, tentacle length index R1 – total length of tentacular stalk and club expressed as a percentage of DML; ClLI, club length index – length of club, measured from proximal-most basal sucker to distal tip of club expressed as a percentage of DML; CIRC, club row count – number of longitudinal rows of suckers across width of club; GLI, gill length index – length of gill expressed as a percentage of DML; GWI, gill width index – greatest width of gill expressed as a percentage of DML; GiLC, gill lamellae count – number of lamellae on outer demibranch, excluding terminal lamella;A SC, sucker counts of arm I – total number of suckers or sucker stalks on arm I; A SC, sucker counts of 1 2 arm II – total number of suckers or sucker stalks on arm II; A SC, sucker counts of arm III – total number of suckers or sucker stalks on arm III; A SC, sucker counts of 3 4 arm IV – total number of suckers or sucker stalks on arm IV; A R1 SC, sucker counts of right arm I – total number of suckers or sucker stalks on right arm I (only in males); ClSI, club sucker index – diameter of largest club sucker expressed as a percentage of DML; A 1 SI, arm I sucker index – diameter of largest sucker on arm I expressed as a percentage of DML; A SI, arm II sucker index – diameter of largest sucker on arm II expressed as a percentage of DML; A SI, arm III sucker index – 2 3 diameter of largest sucker on arm III expressed as a percentage of DML; A SI, arm IV sucker index – diameter of largest sucker on arm IV expressed as a percentage of 4 DML; SpLI, spermatophore length index – length of spermatophore expressed as a percentage of DML; SpWI, spermatophore width index – greatest width of spermatophore expressed as a percentage of spermatophore length; SpRI, sperm reservoir index – length of sperm reservoir expressed as a percentage of spermatophore length; CBI, cement body index – length of cement body expressed as a percentage of spermatophore length.表3 大腹新属新种（Amutatiola macroventosa）雄性标本的测量数据、计数及指数 物种：Amutatiola macroventosa（WAM馆藏） 登记号：F80081、F80083、3091-83、F158244、F275295、F80087、F91362 标本编号：1、2、3、4、5、6、7 模式标本状态：巡航标本、正模（SO1/85）、副模3（SO1/85）、副模2、副模5、N. Sinclair于1983年4月4日采集、副模7、无、无、副模8、无 采集站号：1、2、4、10、1983年4月4日鱼藤酮（Rotenone）采集站、无、无、无 性别：雄性、雄性、雄性、雄性、雄性、雄性、雄性 成熟度：5、5、5、5、4、5、4 注：成熟度4为亚成体——性征已清晰区分，但性腺与附属器官尚未完全发育；成熟度5为成体——雄性内德姆囊（Needham's sac）内含有精荚；雌性的卵巢、缠卵腺与输卵管腺完全发育成熟，卵偶见于输卵管中。 各测量指数定义及结果如下： 1. 背胴长（dorsal mantle length, DML，单位：mm）：从外套膜最前端至外套膜后端的距离，测量值依次为9.8、9.6、9.5、8.8、8.1、8、6.9 2. 腹胴长指数（ventral mantle length index, VMLI）：腹中线处外套膜前缘至外套膜顶端的腹胴长，以背胴长的百分比表示，测量值依次为109.2、93.8、102.1、无、88、116.3、103.6 3. 头长指数（head length index, HLI）：背腕愈合处至背胴膜前端的头部背侧长度，以背胴长的百分比表示，测量值依次为43.9、47.9、40、49.3、59.6、63.8、54.1 4. 头宽指数（head width index, HWI）：眼部水平处头部最大宽度，以背胴长的百分比表示 5. 颈周沟宽指数（nuchal commissure width index, NCI）：颈周沟的宽度，以颈周沟位置处外套膜宽度的百分比表示 6. 胴宽指数（mantle width index, MWI）：外套膜开口处的胴宽，以背胴长的百分比表示，测量值依次为88.8/41.1/96.9、70.8/38.6/91.7、90.5/45.8/87.4、84.2/无/无、87.7/43.7/91.6、101.3/40.2/115、102/50.5/111.4 7. 漏斗长指数（funnel length index, FuLI）：沿腹中线从漏斗前开口至漏斗后缘的漏斗长度，以背胴长的百分比表示，测量值依次为69.4、62.5、70.5、无、81.2、61.3、85.5 8. 漏斗宽指数（funnel width index, FuWI）：漏斗与头部连接处（漏斗锁软骨前方）的漏斗宽度，以背胴长的百分比表示，测量值依次为50、43.8、42.1、34.6、55.1、31.3、58.8 9. 自由漏斗指数（free funnel index, FFuI）：从漏斗前开口至背侧附着于头部处的漏斗长度，以漏斗总长度的百分比表示 10. 鳍长指数（fin length index, FLI）：单个鳍的最大长度，以背胴长的百分比表示，测量值依次为60.3/62.2、66.7/58.3、68.7/53.7、无/无、77.8/54、61.2/63.8、79/64.9 11. 鳍基长指数（fin base length index, FBLI）：鳍附着于外套膜的长度，以背胴长的百分比表示，测量值依次为35.7、30.2、31.6、30.3、29.8、41.3、52.9 12. 鳍宽指数（fin width index, FWI）：单个鳍的最大宽度，以背胴长的百分比表示，测量值依次为46.9、38.5、33.7、59.7、48.1、58.8、58.4 13. 腕I长指数（arm I length index, A1LI）：腕I的长度，以背胴长的百分比表示 14. 腕II长指数（arm II length index, A2LI）：腕II的长度，以背胴长的百分比表示 15. 腕III长指数（arm III length index, A3LI）：腕III的长度，以背胴长的百分比表示 16. 腕IV长指数（arm IV length index, A4LI）：腕IV的长度，以背胴长的百分比表示 17. 腕式（arm formula）：1.2.4.3.1.3.2.4.2.1.4.3.、无、2.1.3.4.2.3.1.4.3.2.1.4. 18. 触腕茎穗总长度指数（tentacle stalk and club total length index, TtLI）：触腕茎与触腕穗的总长度，以背胴长的百分比表示，测量值依次为84.7/无、91.7/161.8、76.8/121.1、102.3/无、72.8/无、100/87.5、88.4/无 19. 触腕穗长指数（club length index, ClLI）：从触腕穗最近端的基部吸盘至触腕穗远端顶端的长度，以背胴长的百分比表示，测量值依次为24.5、21.9、29.5、22.7、32.1、28.8、26.1 20. 触腕穗吸盘列数（club row count, ClRC）：触腕穗宽度方向的纵向吸盘列数，测量值依次为6、6、6、6、6、无、8、无、8、无 21. 鳃长指数（gill length index, GLI）：鳃的长度，以背胴长的百分比表示，测量值依次为42.9/22.4、42.7/16.7、无/无、无/无、40/25.3、33.7/16.3、46.4/18.8 22. 鳃宽指数（gill width index, GWI）：鳃的最大宽度，以背胴长的百分比表示 23. 外鳃瓣鳃片数（gill lamellae count, GiLC，不含末端鳃片）：外鳃瓣上的鳃片数，测量值依次为21、21、无、无、17、16、16 24. 腕I吸盘总数（arm I sucker count, A1SC）：腕I上的吸盘或吸盘柄总数量，测量值依次为20、19、20、14、28、16 25. 腕II吸盘总数（arm II sucker count, A2SC）：腕II上的吸盘或吸盘柄总数量，测量值依次为30/29、30/29、30/27/27、32/21、33/24/24、27/26、28/24/22 26. 腕III吸盘总数（arm III sucker count, A3SC）：腕III上的吸盘或吸盘柄总数量 27. 腕IV吸盘总数（arm IV sucker count, A4SC）：腕IV上的吸盘或吸盘柄总数量 28. 右腕I吸盘总数（right arm I sucker count, AR1SC，仅雄性）：右腕I上的吸盘或吸盘柄总数量，测量值依次为19、18、17、无、12、25、14 29. 触腕穗吸盘指数（club sucker index, ClSI）：触腕穗上最大吸盘的直径，以背胴长的百分比表示，测量值依次为2.4、1.6、1.9、0.6、1.6、2.3、1.9 30. 腕I吸盘指数（arm I sucker index, A1SI）：腕I上最大吸盘的直径，以背胴长的百分比表示，测量值依次为2、1.4/4.7、17.6/9.7、19.8/5.1、18.6/7.3、20.4/17.4、11.9/无 31. 腕II吸盘指数（arm II sucker index, A2SI）：腕II上最大吸盘的直径，以背胴长的百分比表示 32. 腕III吸盘指数（arm III sucker index, A3SI）：腕III上最大吸盘的直径，以背胴长的百分比表示，测量值依次为13.7、8.4、14.5、17.2、20.3、无、20.4 33. 腕IV吸盘指数（arm IV sucker index, A4SI）：腕IV上最大吸盘的直径，以背胴长的百分比表示 34. 精荚长指数（spermatophore length index, SpLI）：精荚的长度，以背胴长的百分比表示，测量值依次为4.1/4.8、a:36.2; b:33.5; c:35.7; d:34.0、38.8、8.1/42.7、5.6/29.3、6.3、无/无、44.8/7.4、无/无 35. 精荚宽指数（spermatophore width index, SpWI）：精荚的最大宽度，以精荚长度的百分比表示，测量值依次为a:6.2; b:6.7; c:5.7; d:6.0、a:33.5; b:35.7; c:34.0; d:43.2、6.7/40.3、5.4/35.5、6.6/37.6、无/无、7.0/27.9、无/无 36. 储精囊指数（sperm reservoir index, SpRI）：储精囊的长度，以精荚长度的百分比表示 37. 黏固体指数（cement body index, CBI）：黏固体的长度，以精荚长度的百分比表示，测量值依次为a:20.3; b:22.0; c:19.1; d:20.0、19.4、18.5、19.4、无、10.9、无