The relative contribution (%) of the predictor variables to the BRT interaction models over the local scale.

The predictor variables were Beach, mean grain size (MGS), beach slope, and sediment shear strength (SS). The predictor variables with the highest contribution described best the recorded observations of the main beach species. Beach species included in the final model: Bathyporeia pelagica, Donax trunculus, Eurydice affinis, Eurydic pulchra, Pontocrates arenarius, Scolelepis squamata, Sphaeroma rugicauda, Talorchestia brito and Talitrus saltator.