TABLE 1 in Description of a new species of the earthworm genus Moniligaster Perrier, 1872 (Clitellata, Moniligastridae) from the Nilgiri Biosphere Reserve of Tamil Nadu State, India

TABLE 1. Comparison of the characters of M. gatesi Ahmed & Narayanan, sp. nov., with other species of the ‘ gravelyi ’ group.
CharacterM. gravelyi Stephenson, 19151,2M. troyi Jamieson, 19773M. bahli Narayanan & Julka,20214M. blakemorei Narayanan & Julka, 20214,5M. keralensis Narayanan & Julka, 20214,5M. julkai Narayanan & Paliwal, 20225M. gatesi Ahmed & Narayanan, sp. nov.Length130–174 mm60–63 mm75–134 mm81–136 mm58–155 mm156–238 mm87–160 mmDiameter5–7 mm3.5–3.7 mm4.5–7 mm3.5–7 mm3–6 mm6.5–8.5 mm5–7 mmSegments139–189123–153135–200183–278109–201237–342116–181Spermathecal pore locationbetween bc setal lines, median to setae cin c setal linebetween bc, median to setae cclose to cd setal linesaligned between cd setal linesaligned at cd setal linesbetween bc setal lines, nearer to cClitellumannular, on segments ¼9– ½14 (= ¾4)annular, on segments ½9–14 (=5½)annular, on segments 10–13 (=4)annular, on segments 10–13 (=4)annular, on segments 10–13 (=4)annular, on segments 10–½14 (=4½)annular, on segments ⅓9–14 (=5⅓)Secondary male aperture or male pore locationaligned between bc setal lines, nearer to bon evaginated long cylindrical pseudopenes, at bc setal lines, lateral to blateral to seta b linesslightly lateral to b setal lineslightly lateral to b setal lineslateral to b setal linebetween bc setal lines, nearer to bFemale pore locationat b setal line?minute, at b setal lineminute, at b setal lineminute, slightly median to b setal lineminute, at b setal line?b setal lineSpermathecaatrial gland single, on each side; duct of the gland c. 1/7 the length of common atrial duct; ampulla ovoidatrial glands paired on each side; duct of each gland c. 1/4 the length of common atrial duct; ampulla elongate-ovoidatrial glands paired on each side; duct of each gland c. 1/5 the length of common atrial duct; ampulla ovoidatrial glands paired on each side; duct of each gland c. four times the length of common atrial duct; ampulla ovoidalatrial glands paired on each side; duct of each gland about as long as common atrial duct; ampulla elongate-ovoidatrial glands paired on each side; duct of each gland c. five times the length of common atrial duct; ampulla ovoidalatrial glands paired on each side; duct of the gland c. 1/6 the length of common atrial duct; ampulla pear-shapedProstatesstrap-shapedclub-shapedstrap-shapedbluntly club-shapedtubulartubularstrap-shapedProstate capsuletubular with deeply incised margins and highly nodulated surface(?)tubular with smooth margins and a few nodulations at ental endclub-shaped with smooth marginstubular with smooth marginstubular with smooth marginstubular, surface with shallow nodulations, margins nodulatedLength of prostatic ductjust over 1/4 the length of gland plus ductc.1/3 the length of gland plus ductc. 1/7 the length of gland plus ductc. 1/3 the length of gland plus ductc. 1/9 the length of gland plus ductc. 1/3 the length of gland plus ductc. 1/8 the length of gland plus ductVas deferenssegments 9–10, 11, enters prostate sub-entallysegment 9, enters prostate sub-ectallysegments 9–12(–16), enters prostate sub-entallysegments 9–11(– 12), enters prostate near entallysegments 9–10, enters prostate sub-ectallysegments 9–12(–13), enters prostate sub-ectallysegments 9–10, enters prostate sub-entallyGizzards4–5 (in segments 12–18)3 (in segments 13–15)3–4 (in segments 13–18)2 (in segments 12–16)3–6(in segments 14–20)2–3(in segments 12–16)4–5 (in segments 13–18)Intestinal originin segment 25±1in segment 27in segment 25±1in segment 27±1in segment 28±1in segment 27in segment 26±1Ovisacsextending to segments 13–14extending to segment 13extending to segment 20extending to segment 13extending to segments 12–16extending to segments 13–14extending to segments 13–16
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Data from: 1 Stephenson (1915); 2 Narayanan et al. (2023b); 3 Jamieson (1977); 4 Narayanan et al. (2021a); 5 Narayanan et al. (2022).