Diversity and Systematics of Schizomavella Species (Bryozoa: Bitectiporidae) from the Bathyal NE Atlantic

Eight NE Atlantic and Mediterranean species, which were originally assigned to the genus Schizoporella (Family Schizoporellidae) when introduced, are redescribed and stabilized by typification. Seven of these species are transferred to the bitectiporid genus Schizomavella: S. fischeri, S. glebula, S. neptuni, S. obsoleta, S. richardi, S. triaviculata, and S. triaviculata var. paucimandibulata, which is here raised to species rank. The eighth species, Schizoporella fayalensis, is transferred to the lanceoporid genus Stephanotheca. Schizomavella obsoleta and S. glebula are considered junior subjective synonyms of S. fischeri and S. richardi, respectively. Two new species are described: Schizomavella rectangularis n. sp. from the Strait of Gibraltar, and Schizomavella phterocopa n. sp. from the Great Meteor Bank. A new subgenus, Calvetomavella n. subgen. is established as a result of a phylogenetic analysis based on morphological characters; it includes S. neptuni, S. triaviculata, S. paucimandibulata and S. phterocopa n. sp., together with Schizomavella discoidea and Schizomavella noronhai. The rest of the species remain in the nominotypical subgenus Schizomavella.8种分布于东北大西洋及地中海的物种，在最初发表时被归入裂孔苔虫科（Schizoporellidae）的裂孔苔虫属（Schizoporella），现通过模式标定对其进行重新描述并完成分类稳定化。其中7个物种被归入双节苔虫科（Bitectiporidae）的裂爪苔虫属（Schizomavella），分别为S. fischeri、S. glebula、S. neptuni、S. obsoleta、S. richardi、S. triaviculata，以及原作为S. triaviculata变种的S. triaviculata var. paucimandibulata，本文将其提升为独立物种。第8个物种——Schizoporella fayalensis，被归入尖孔苔虫科（Lanceoporidae）的斯蒂芬苔虫属（Stephanotheca）。裂爪苔虫属的S. obsoleta与S. glebula分别被视为S. fischeri与S. richardi的次级主观同物异名。本文还描述了2个新物种：产自直布罗陀海峡的Schizomavella rectangularis n. sp.，以及产自大流星海山的Schizomavella phterocopa n. sp.。基于形态特征的系统发育分析结果表明，本文建立一新亚属Calvetomavella n. subgen.，该亚属包含S. neptuni、S. triaviculata、S. paucimandibulata与S. phterocopa n. sp.，以及Schizomavella discoidea与Schizomavella noronhai。其余物种仍归入裂爪苔虫属（Schizomavella）的指名亚属。