Table 3 in Deepest known novel species of the genus Ophiuroglypha Hertz, 1927 (Echinodermata: Ophiuroidea) from the central rift zone, Philippine Sea

Table 3 (continued on next two pages). Tabular key to all species of the genus Ophiuroglypha Hertz, 1927, compiled from the literature. For convenience, the second oral tentacle pore is here labelled as the 1 st pore, because it is the first visible pore along the arm, the actual first pore being placed laterally in the oral plates (= jaw) and both ventral tentacle scales and adoral shield spines in second oral tentacle pore labelled as tentacle scales. Abbreviations: ASE = arm segment; DAP = dorsal arm plate; LOP = lateral oral papillae; TS = tentacle scale; VAP = ventral arm plate; VMT = ventralmost tooth. Speciesarm spinesRadial shieldsDorsal discOral frameTentacle scalesVAPDAPReferencesOphiuroglypha aequatorisup to 3, evenlylarge, wide,scales small4–5 LOP, 11st pore 4–5 TS atcontiguous, 1stwider than long,Hertz (1927)Hertz, 1927spaced, ½ ASEseparatedin center,spiniform VMTeach side; 2nd poreoctagonal, distallyfan-shaped,long, middle spineproximally butlarger distally,4–5 TStriangularcontiguousglassy hookconnected at distalcentrodorsal plateendsrecognizableOphiuroglypha ambiguaup to 4, short, blunt, small, triangular,moderate in size,4–5 LOP, 11st pore 4–5 TS at1st large, diamond-fan-shaped,Lyman (1878)(Lyman, 1878)tooth-like, closeseparated by 1thick and swollenspiniform VMTeach side; 2nd poreshaped, thencontiguoustogetherlarge and 2–3 smallirregular scales5– TS, distally 1 TS pentagonal,scalescontiguousproximally, butseparated distallyOphiuroglypha arntziup to 3, small,elevated from disc,irregular scales,4–5 LOP, 1–21st pore 4–5 TS at1st –3rd contiguous,contiguousManso (2010)(Manso, 2010)thick, middle spineseparated by tworadial primaryspiniform VMTeach side; 2nd porethen separated;hooklarge scalesplates rounded,5–7 TSwider than longsmall scalesbetween radialprimary plates andcentrodorsal plateOphiuroglypha brevispinosaup to 6–7, shortequal in size tomoderately large4 LOP, 3 thick,1st pore 5–6 TS atcontiguouslonger than broad,Smith (1876), Hertz(H.L. Clark, 1915)disc scales, distally5–6 radial primaryconical VMTeach side; 2nd–3rdcontiguous(1927), H.L. Clarkcontiguousplates, irregular discpores 5–6 TS(1915)scalesOphiuroglypha clemensUp to 3, small, andtriangular shapedspaced irregular4 LOP, blunt,1st–2nd pores 5–6separatedcontiguous(Koehler 1904)(Koehler, 1904)unevenly spacedwith curved edge,disc scales,pointed VMTTS at each side; 3rdcontiguousradial primarypore 3–4 TS, thenplates cannot beTS fewerdistinguishedOphiuroglypha cariniferaup to 3, small,separated by twosix radial4–5 LOP, 1–21st pore 5–6 TS at1st–3rd contiguouscontiguousKoehler (1901)(Koehler, 1901)conicalscalesprimary platespiniform VMTeach side; 2nd porethen separated,rosette distinct,5–6 TS, 3rd pore 4–5 1st triangular, 2ndcentrodorsalTS, then 3 TS andquadrangular, thenroundedfinally 2 or 1 TStriangularOphiuroglypha costataup to 3, small,long, rhomboidal,few, large, rounded4-5 LOP, 11st pore 5 TS at1st broad triangular,fan-shaped,Lyman (1878,(Lyman, 1878)blunt, peg-like,slightly curvedscales, separatedspiniform VMTeach side; 2nd porebeyond pentagonal,contiguous1882), Bell (1905),evenly spacededge, separated5 radial primary5 TS, 3rd pore 3 TS,separatedHertz (1927),plates, centrodorsaldistally reduce intoMortensen (1933),rounded1 TSA.M. Clark (1952), Olbers et al. (2019)Ophiuroglypha ossiculataup to 9, small,small, irregularirregular plates,5 LOP, 1 spiniform1st pore 5–6 TS at1st fan-shaped, 2ndrectangular, short,Koehler (1908)(Koehler, 1908)papilliformmore or lesswell-spacedVMTeach side; 2nd poretrapezoidal, beyondwide, contiguoustriangular, separated rounded primary5–6 TS, 3rd andtriangular, thenby 1–2 large scalesplates4th pore 4–5 TS,separated5th pore 3–4 TS,beyond 3 TSOphiuroglypha planaup to 3, upper one 1 small, separated bycoarser or thinner4-5 LOP, 11st pore 6–9 TS at1st polygonal, 2ndfan-shaped,Lütken &(Lütken & Mortensen, 1899)ASE length1 or more scalesoverlapping scales,spiniform VMTeach side; 2nd porepentagonal, beyondcontiguousMortensen (1899)primary plates6–9 TS, 3rd pore 5–7 5th separatedrather conspicuousTS, distally reduceinto 1 TSOphiuroglypha rugosaup to 3, evenlylonger thanlarge, swollen4 LOP, 1 spiniform1st pore 4–5 TS atseparated, 1stlong, wedge-Lyman (1878)(Lyman, 1878)spaced, short, bluntwide, proximallyscales with deepVMTeach side; 2nd–3rdtriangular, beyondshaped, contiguousseparated by largefurrow, 5 radialpores 4 TS, 4th porepentagonalscales, distallyprimary plates and3 TS, 5th pore 1–3contiguousone central plate,TS, beyond 1 TSseparated by smallirregular scalesOphiuroglypha schmidtottiup to 3, evenlytriangular or oval,thick disc scales,3–6 LOP, all broad,1st pore 5 TS at each separated, 1st bellcontiguousHertz (1927),Hertz, 1927spaced, short,distally contiguousirregular size,1–3 broad VMTside; 2nd pore 5 TS,shaped, beyond fan-Paterson (1985),blunt, one hookeddistinct primarydecreasing to 1 TSshapedOlbers et al. (2019)upwardsrosette with largecentrodorsal plateOphiuroglypha scombup to 3, small, ½small, pear-shaped,large irregular5–6 LOP, 1–31st pore 7 TS at each separated, 1st bell-contiguous, distallyPaterson (1985)(Paterson, 1985)ASE lengthdistally contiguousscales, largespiniform VMTside; 2nd pore 6–9shaped, beyondseparatedor separatedcentro-dorsal plate,TS, 3rd pore 7–11fan-shapedprimary platesTS, 4th pore 4–7 TSdistinctOphiuroglypha tumidaup to 3, short, upper longer than wide,irregular, medium3–5 LOP, 1mouth pore 6–9 TSBell-shaped,Fan-shaped,Mortensen (1933),(Mortensen, 1933)one slightly longeroval, separated bysize scales, primaryspiniform VMTat each side; 1st pore separatedseparatedOlbers et al. (2019)one scalerosette distinctup to 9 TS, distallyreduce into 1 TSOphiuroglypha verrucosaup to 3, two near TS smaller than otherdistinct spaced5–6 LOP, 1–21st pore 6–11 TS atseparated, widercontiguous, widerMcKnight (2003)(McKnight, 2003)scales, broad,scales, irregularspiniform VMTeach side; 2nd–6ththan longthan longseparatedin shape, roundedpores 3–5 TS,or pentagonalbeyond 1 TScentrodorsal plate表3（续于后两页）。本文基于文献汇编了蛇形海蛇尾属（Ophiuroglypha Hertz, 1927）所有物种的分类检索表。为便于表述，本文将第二口触手孔标记为第1触手孔——因其为腕部首个可见的触手孔，而真正的第1触手孔位于口板（即颚）的侧面；同时将第二口触手孔的腹侧触手鳞与口盾棘均归类为触手鳞。 缩写说明： ASE = 腕节（arm segment）； DAP = 背腕板（dorsal arm plate）； LOP = 侧口乳头（lateral oral papillae）； TS = 触手鳞（tentacle scale）； VAP = 腹腕板（ventral arm plate）； VMT = 最腹面齿（ventralmost tooth）。 以下为各物种的分类性状表格： | 物种 | 腕棘 | 辐盾 | 盘背结构 | 口部构造 | 触手鳞 | 腹腕板 | 背腕板 | 参考文献 | | --- | --- | --- | --- | --- | --- | --- | --- | --- | | 赤道蛇形海蛇尾（Ophiuroglypha aequatoris） | 最多3枚，均匀分布，长度为腕节的1/2 | 大型、宽幅，中央分离，远端连续 | 呈八边形，远端呈扇形，宽大于长，中间棘远端更大，玻璃状钩突于远端连接处清晰可辨 | 鳞片小型，中央处为纺缍形最腹面齿 | 第1触手孔：每侧具4~5枚侧口乳头，4~5枚触手鳞；第2触手孔：近端4~5枚触手鳞，远端更大 | 相邻、长大于宽 | 赫茨（1927） | | 歧义蛇形海蛇尾（Ophiuroglypha ambigua） | 最多4枚，短而钝，呈齿状，闭合时紧贴 | 小型、三角形，中等大小，厚实且肿胀 | 扇形，纺缍形最腹面齿 | 第1触手孔：每侧具4~5枚侧口乳头，4~5枚触手鳞；第2触手孔：近端2~3枚小型鳞片，远端1枚鳞片 | 第1枚大型，呈菱形，后变为不规则鳞片，近端连续、远端分离 | 相邻 | 莱曼（1878） | | 阿恩茨蛇形海蛇尾（Ophiuroglypha arntzi） | 最多3枚，小型，中间棘厚实 | 从盘上抬起，具不规则鳞片，2枚辐盾原板，中央板圆形，辐盾原板与中央板之间具小型鳞片 | 不规则鳞片，间距2枚大型鳞片 | 第1触手孔：每侧具4~5枚侧口乳头，1~2枚纺缍形最腹面齿；第2触手孔：近端4~5枚触手鳞，后逐渐分离 | 相邻，后分离 | 曼索（2010） | | 短棘蛇形海蛇尾（Ophiuroglypha brevispinosa） | 最多6~7枚，短，与盘鳞大小一致 | 盘鳞大小中等，远端具4~5枚辐盾原板，不规则盘鳞 | 圆锥形最腹面齿 | 第1触手孔：每侧具4枚侧口乳头，3枚厚实结构；第2~3触手孔：每侧5~6枚触手鳞 | 长大于宽，相邻 | 史密斯（1876）、赫茨（1927）、H.L.克拉克（1915） | | 克莱门斯蛇形海蛇尾（Ophiuroglypha clemens） | 最多3枚，小型，分布不均 | 三角形，边缘弯曲，盘鳞具不规则间隔 | 具尖齿状最腹面齿 | 第1~2触手孔：每侧具4枚侧口乳头，5~6枚触手鳞；第3触手孔：具3~4枚触手鳞，后续更少 | 相邻，无法区分辐盾原板 | 科勒（1904） | | 龙骨蛇形海蛇尾（Ophiuroglypha carinifera） | 最多3枚，小型，圆锥形 | 间距2枚大型鳞片，6枚辐盾原板 | 具纺缍形最腹面齿 | 第1触手孔：每侧具4~5枚侧口乳头，1~2枚；第2触手孔：每侧5~6枚触手鳞，后分离；第3触手孔：4~5枚触手鳞，后续3~2~1枚触手鳞 | 第1枚三角形，第2枚四边形，后变为三角形，具清晰的花冠状中央板 | 相邻 | 科勒（1901） | | 肋纹蛇形海蛇尾（Ophiuroglypha costata） | 最多3枚，小型，短而钝，呈桩状，略弯曲 | 长菱形，具少量大型圆形鳞片，间距均匀，5枚辐盾原板，中央板圆形 | 纺缍形最腹面齿 | 第1触手孔：每侧具4~5枚侧口乳头，5枚触手鳞；第2触手孔：5枚触手鳞；第3触手孔：3枚触手鳞，远端缩减为1枚 | 宽三角形，远端呈扇形，相邻 | 莱曼（1878、1882）、贝尔（1905）、赫茨（1927）、莫滕森（1933）、A.M.克拉克（1952）、奥尔伯斯等（2019） | | 骨鳞蛇形海蛇尾（Ophiuroglypha ossiculata） | 最多9枚，小型，乳突状 | 小型、不规则，分布均匀 | 具纺缍形最腹面齿 | 第1触手孔：每侧具5枚侧口乳头；第2触手孔：5~6枚触手鳞；第3~4触手孔：4~5枚触手鳞；第5触手孔：3~4枚触手鳞，远端超过3枚 | 第1枚扇形，第2枚梯形，后变为三角形，间距1~2枚大型鳞片，圆形原板 | 分离 | 科勒（1908） | | 扁平蛇形海蛇尾（Ophiuroglypha plana） | 最多3枚，上方1枚长度为腕节的1倍或更多 | 小型，被1枚或多枚鳞片覆盖，重叠鳞片、粗或细的鳞片 | 具纺缍形最腹面齿 | 第1触手孔：每侧具4~5枚侧口乳头；第2触手孔：6~9枚触手鳞；第3触手孔：5~7枚触手鳞，远端缩减为1枚 | 第1枚多边形，第2枚五边形，后分离，具显著原板 | 扇形 | 吕特肯&莫滕森（1899） | | 皱皮蛇形海蛇尾（Ophiuroglypha rugosa） | 最多3枚，均匀分布，短而钝 | 长大于宽，近端具深沟，5枚辐盾原板与1枚中央板 | 具深沟的大型肿胀鳞片 | 第1触手孔：每侧具4枚侧口乳头，1枚纺缍形最腹面齿；第2~3触手孔：每侧4枚触手鳞；第4触手孔：3枚触手鳞；第5触手孔：1~3枚触手鳞，远端超过1枚 | 分离，第1枚长楔形，相邻，后被大型鳞片分离 | 分离，被小型鳞片分隔 | 莱曼（1878） | | 施密特蛇形海蛇尾（Ophiuroglypha schmidtotti） | 最多3枚，均匀分布，短而钝，具1个钩状突 | 三角形或椭圆形，远端连续，具大小不规则的明显花冠状中央板 | 厚实的盘鳞 | 第1触手孔：每侧具3~6枚侧口乳头，1~3枚宽幅最腹面齿；第2触手孔：每侧5枚触手鳞，向上逐渐减少至1枚 | 分离，第1枚钟形，后变为扇形 | 赫茨（1927）、帕特森（1985）、奥尔伯斯等（2019） | | 鲭蛇形海蛇尾（Ophiuroglypha scomb） | 最多3枚，小型，长度为腕节的1/2，远端连续 | 小型、梨形，远端连续 | 大型不规则鳞片 | 第1触手孔：每侧具5~6枚侧口乳头，1~3枚纺缍形最腹面齿；第2触手孔：6~9枚触手鳞；第3触手孔：7~11枚触手鳞；第4触手孔：4~7枚触手鳞 | 分离，第1枚钟形，后变为扇形，具明显中央板与辐盾原板 | 相邻，远端分离 | 帕特森（1985） | | 肿胀蛇形海蛇尾（Ophiuroglypha tumida） | 最多3枚，短，上方1枚长大于宽 | 椭圆形，被1枚鳞片分隔，中等大小的不规则鳞片，具显著花冠状结构 | 具纺缍形最腹面齿 | 第1触手孔：每侧具3~5枚侧口乳头，1枚最腹面齿；第2触手孔：6~9枚触手鳞，每侧；第1触手孔分离，远端缩减为1枚 | 钟形，分离 | 莫滕森（1933）、奥尔伯斯等（2019） | | 疣突蛇形海蛇尾（Ophiuroglypha verrucosa） | 最多3枚，2枚靠近，鳞片小于其他鳞片 | 宽大于长，不规则形状，圆形或五边形，分离 | 具1~2枚宽幅最腹面齿 | 第1触手孔：每侧具5~6枚侧口乳头，1~2枚；第2~6触手孔：3~5枚触手鳞，远端超过1枚 | 分离，宽大于长，相邻 | 麦基特里克（2003） |