TABLE. 2 in Three Novel Species of Earthworms of Genus Metaphire Sims and Easton, 1972 from Manipur, India

TABLE. 2. List of comparative traits of novel species with related species.
S. No.Morphological charactersMetaphire manipurensis sp. nov.M. houlleti (Perrier 1872)M. harrietensis (Stephenson 1925)M. scitula (Gates1936)M. umbraticola (Gates 1932)M. quadrigemina (Gates 1932)M. perichaeta (Beddard 1900)M. virgo (Beddard 1900)M. acampanulata Nguyen, Ly, Lam, Nguyen & Nguyen, 20221Body length (mm)80–21075–902001001156416015277–1982Diameter (mm)4.5–4.83.5–3.711.5564554.03–6.913Segment number61–12382–93123?12511511812956–1444First dorsal pore11/129/1012/1312/1312/1312/1312/1311/1211/12 or 12/135Male pore setae12–149–122812–148612128–256Male porein copulatory pouch with transverse slit like opening male genital region expanded laterallyin copulatory pouch with C-shaped openingthe male pores are on conical papillae on xviiiat the centre of a smooth circular disc, deeply retracted into a parietal invagination, with a short transverse slit-like apertureapertures are wide slits in the setal circle of xviii with wrinkled margins.apertures are transverse slits in the setal circle of xviii with antero posterior lips in contact with each otherconspicuous orifices separated from each other by a fairly wide intervalminute and on a porophore within a copulatory chamber opening to the exterior through a transversely slit-like equatorial apertureinside copulatory pouches in xviii, and the openings slit or round shaped.7Spermathecaeampulla obovate or ovate, duct stout, conical, muscular with flat top, long stalked diverticulum terminating in a zig-zag looped swollen receptacle.ampulla oval, with relatively larger swollen spermathecal duct; diverticulum enlarged with looped end.ampulla saclike, shortly ovoid in shape, duct moderately stout not shining, diverticulum long and thin, and irregularly twisted.spermathecae small, duct almost confined to the parietes, diverti culum long looped or zigzag.ampulla is longer than the duct, diverticulum composed of a thick-walled stalk.duct is short, diverticulum bent into a series of zigzag loops.ampulla pear shape, diverticulum zigzagsmall, diverticulum tubularampulla mushroom-shaped with longitudinal grooves on the surface, L-shaped diverticulum.8Genital marking gland2 stalked spherical or mushroom shaped in vii, viii, and xi, buried in parietes.single stalked, connecting to spermathecal ducts. Sometimes, 2–3 sessile in copulatory pouchvery numerous corresponding to the pitted areas stalked.absentabsentabsentabsentone finely acinous, 2–4 with short stalks passing to markings within the chamber2–4 stalked, attached to each ampulla duct. 3–4 in wall of each copulatory pouch in segment xviii.9Lateral intestinal caecaIn xxvii, and extends upto xxiixxvii–xxvxxvii–?xxvii–?xxvii–xxivxxvii into xxvxxvii–xxivxxvii–xxvxxvii–xxiv10Intestinal originxivxvxvxvixvxvixvixvxv
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表2. 新种与近缘物种的比较性状列表
1. 体长（mm）：曼普尔远盲蚓（Metaphire manipurensis sp. nov.）80–210；胡莱特远盲蚓（M. houlleti (Perrier 1872)）75–90；哈里特远盲蚓（M. harrietensis (Stephenson 1925)）200；秀丽远盲蚓（M. scitula (Gates 1936)）100；阴地远盲蚓（M. umbraticola (Gates 1932)）115；四孔远盲蚓（M. quadrigemina (Gates 1932)）64；围毛远盲蚓（M. perichaeta (Beddard 1900)）160；处女远盲蚓（M. virgo (Beddard 1900)）152；无环远盲蚓（M. acampanulata Nguyen, Ly, Lam, Nguyen & Nguyen, 2022）77–198
2. 体宽（mm）：曼普尔远盲蚓4.5–4.8；胡莱特远盲蚓3.5–3.7；哈里特远盲蚓11.5；秀丽远盲蚓5；阴地远盲蚓6；四孔远盲蚓4；围毛远盲蚓5；处女远盲蚓4.0–6.9；无环远盲蚓13
3. 体节数：曼普尔远盲蚓61–123；胡莱特远盲蚓82–93；哈里特远盲蚓123？；秀丽远盲蚓？；阴地远盲蚓125；四孔远盲蚓115；围毛远盲蚓118；处女远盲蚓129；无环远盲蚓56–144
4. 首个背孔位置：曼普尔远盲蚓11/12；胡莱特远盲蚓9/10；哈里特远盲蚓12/13；秀丽远盲蚓12/13；阴地远盲蚓12/13；四孔远盲蚓12/13；围毛远盲蚓12/13；处女远盲蚓11/12；无环远盲蚓11/12或12/13
5. 雄性刚毛孔数：曼普尔远盲蚓12–14；胡莱特远盲蚓9–12；哈里特远盲蚓28；秀丽远盲蚓12；阴地远盲蚓12–14；四孔远盲蚓8；围毛远盲蚓6；处女远盲蚓12；无环远盲蚓8–25
6. 雄性孔：曼普尔远盲蚓位于交配囊内，开口呈横裂隙状；胡莱特远盲蚓雄性生殖区向两侧扩展；哈里特远盲蚓位于交配囊内，开口呈C形；秀丽远盲蚓雄性孔位于第xviii体节的锥形乳突上；阴地远盲蚓位于光滑圆形盘中央，深陷于体壁内陷中，具短横裂隙状开口；四孔远盲蚓开口为第xviii体节刚毛环内的宽裂隙，边缘具褶皱；围毛远盲蚓开口为第xviii体节刚毛环内的横裂隙，前后唇相互贴合；处女远盲蚓雄性孔显著，彼此间隔较宽；无环远盲蚓位于第xviii体节的交配囊内，开口呈裂隙状或圆形
7. 受精囊（spermathecae）：曼普尔远盲蚓囊体倒卵形或卵形，导管粗壮、锥形且具肌肉质平顶，具长柄盲管，盲管末端为呈Z字形盘绕的膨大端囊；胡莱特远盲蚓囊体卵形，受精囊导管相对更膨大；盲管膨大，末端盘绕；哈里特远盲蚓囊体囊状，呈短卵形，导管中等粗壮无光泽，盲管细长且不规则扭曲；秀丽远盲蚓受精囊微小，导管几乎附着于体壁，盲管长且盘绕或呈Z字形；阴地远盲蚓囊体长于导管，盲管由厚壁柄构成；四孔远盲蚓导管短，盲管盘绕成一系列Z字形环；围毛远盲蚓囊体呈梨形，盲管呈Z字形；处女远盲蚓受精囊微小，盲管呈管状；无环远盲蚓囊体呈蘑菇形，表面具纵沟，盲管呈L形
8. 生殖腺斑：曼普尔远盲蚓第vii、viii、xi体节具2个具柄球形或蘑菇形腺体，埋于体壁内；胡莱特远盲蚓单柄状，连接于受精囊导管，有时具2–3个无柄结构位于交配囊内；哈里特远盲蚓数量极多，对应具凹陷区域；秀丽远盲蚓缺失；阴地远盲蚓缺失；四孔远盲蚓缺失；围毛远盲蚓缺失；处女远盲蚓具1个细腺泡状结构，2–4个具短柄，连接于腺室内的斑痕；无环远盲蚓2–4个具柄，附着于每个囊体导管，第xviii体节的每个交配囊壁内具3–4个
9. 侧肠盲囊：曼普尔远盲蚓位于第xxvii体节，延伸至第xxii体节；胡莱特远盲蚓第xvii–xxv体节；哈里特远盲蚓第xxvii–？体节；秀丽远盲蚓第xxvii–？体节；阴地远盲蚓第xxvii–xxiv体节；四孔远盲蚓第xxvii体节延伸至第xxv体节；围毛远盲蚓第xxvii–xxiv体节；处女远盲蚓第xxvii–xxv体节；无环远盲蚓第xxvii–xxiv体节
10. 肠起始节段：曼普尔远盲蚓第xiv体节；胡莱特远盲蚓第xv体节；哈里特远盲蚓第xv体节；秀丽远盲蚓第xv体节；阴地远盲蚓第xv体节；四孔远盲蚓第xv体节；围毛远盲蚓第xv体节；处女远盲蚓第xv体节；无环远盲蚓第xv体节
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