A new semionotiform actinopterygian fish from the Mesozoic of Spain and its phylogenetic implications

<i>Camerichthys lunae</i> gen. et sp. nov. is a semionotiform actinopterygian fish described from the newly reported locality of San Andrés de San Pedro (province of Soria, Spain). The material, an articulated, partial three-dimensional specimen, was unearthed from the Tithonian–Berriasian beds of the Matute Formation (Tera Group) in the continental Cameros Basin. The new taxon, <i>Camerichthys lunae</i> gen. et sp. nov., differs from other ginglymodians in presenting a unique combination of characters plus: presence of a suprapreopercular bone; antorbital intermediate in depth in relation to the other anterior infraorbitals; and infraorbital placed in the anterior border of the orbit subdivided into two portions (three uniquely derived characters). The results of the cladistic analysis show that the new genus from Soria cannot be assigned to any of the three semionotiform families, Callipurbeckiidae, Macrosemiidae or Semionotidae. It is considered Semionotiformes <i>incertae sedis</i>. In the more resolutive analysis, with ordered characters, <i>Camerichthys lunae</i> gen. et sp. nov. appears as the sister group of the clade formed by the families Callipurbeckiidae plus Macrosemiidae. The effects of the new genus on the phylogeny of these two families and <i>Dapedium</i>, <i>Semionotus</i> and <i>Paralepidotus</i>, are discussed. <i>Camerichthys</i> gen. nov. is the first ginglymodian genus endemic to the Iberian Peninsula.http://zoobank.org/urn:lsid:zoobank.org:pub:BBD15A41-5BFC-4BFF-AD40-ECD1891953F1

新月卡梅里鱼（Camerichthys lunae）新属新种是一类半椎鱼目辐鳍鱼类，其化石发现于西班牙索里亚省新报道的圣安德烈斯德圣佩德罗化石产地。该标本为关节连接完好的部分三维化石标本，出土于陆相卡梅罗斯盆地内马图特组（泰拉群）的提通阶-贝里亚斯阶地层中。
该新分类单元具备独特的特征组合，可与其他硬鳞总类（ginglymodians）相区分：存在前鳃盖骨上骨（suprapreopercular bone）；眶前骨（antorbital）的深度介于其余前部眶下骨（infraorbital）之间；位于眼眶前缘的眶下骨（infraorbital）分为两部分，共计3个独有衍征。分支系统学分析（cladistic analysis）结果显示，这一产自索里亚省的新属无法归入半椎鱼目现有的三个科——卡利珀贝克鱼科（Callipurbeckiidae）、大鳞半椎鱼科（Macrosemiidae）与半椎鱼科（Semionotidae），因此被归为半椎鱼目分类位置未定类群（incertae sedis）。
在采用有序性状的高分辨率分支分析（resolutive analysis）中，新月卡梅里鱼新属新种作为由卡利珀贝克鱼科与大鳞半椎鱼科共同构成的演化支（clade）的姊妹群（sister group）出现。本文还探讨了该新属对上述两科以及戴普鱼属（Dapedium）、半椎鱼属（Semionotus）与副鳞鱼属（Paralepidotus）系统发育关系的影响。新月卡梅里鱼新属是首个仅分布于伊比利亚半岛的硬鳞总类属。
http://zoobank.org/urn:lsid:zoobank.org:pub:BBD15A41-5BFC-4BFF-AD40-ECD1891953F1